心叶三脉蕨(新属、新组合)及大羽羊齿类的分类位置

TRINERVIOPTERIS CARDIOPHYLLA (ZHU ET GENG) ZHU GEN. ET COMB. NOV. AND THE CLASSIFICATION OF GIGANTOPTERIDES

报道大羽羊齿类中的一真蕨植物新属和新组合。标本采自福建将乐县晚古生代的“龙潭组”。曾以脉序介于重网状和单网状之间，鉴定为心叶单网羊齿（新种）（Ｇｉｇａｎｔｏｎｏｃｌｅａ ｃａｒｄｉｏ－ｐｈｙｌｌａ Ｚｈｕ ｅｔＧｅｎｇ）。重新研究后，根据其三出脉和不完全的重网状脉式，又发现了其孢子囊群，从营养叶看，可归入大羽羊齿类，介于大羽羊齿和单网羊齿之间，从孢子囊群看，则肯定是真蕨，重新命名为心叶三脉蕨（新属、新组合）［Ｔｒｉｎｅｒｖｉｏｐｔｅｒｉｓ ｃａｒｄｉｏｐｈｙｌｌａ （Ｚｈｕ ｅｔ Ｇｅｎｇ）Ｚｈｕ ｇｅｎ．ｅｔ ｃｏｍ ｂ．ｎｏｖ．］。与之同时发现的其它植物有：Ｓｐｈｅｎｏｐｈｙｌｌｕｍ ｅｍ ａｒｇｉｎａｔｕｍ 、Ｂｏｗ －ｍ ａｎｉｔｅｓ ｌａｘｕｓ、Ｌｏｂａｔａｎｎｕｌａｒｉａ ｌｉｎｇｕｌａｔａ、Ｃｈａｎｓｉｔｈｅｃａ ｐａｌａｅｏｓｉｌｖａｎａ、Ｃｈ． ｃｆ． ｋｉｄｓｔｏｎｉｉ、Ｐｅｃｏｐｔｅｒｉｓ ａｎｄｅｒｓｓｏｎｉｉ、Ｐ．ｅｃｈｉｎａｔａ、Ｐ．？ ｔａｉｙｕａｎｅｎｓｉｓ、Ｐ．ｕｎｉｔａ、Ｄａｎａｅｉｔｅｓ ｍ ｉｒａｂｉｌｉｓ、Ｆａｓｃｉｐｔｅｒｉｓｄｅｎｓａｔａ、Ｃｌａｄｏｐｈｌｅｂｉｓ ？ ｐｅｒｍ ｉｃａ。

A new fossil genus and combination of gigantopterides is reported. The specimen was collected from the Late Paleozoic “Longtan Formation” of Jiangle county, Fujian Province of China. It was originally determined to the genus Gigantonoclea and named as G.cardiophylla Zhu et Geng by one of the present authors and his colleague some times ago. In view of its venation being trinervious and that the incompletely duplicated net like veins are close to those of gigantopterides and the leaf bears sori on its back. Upon reconsideration it was renamed by the authors as Trinerviopteris cardiophylla (Zhu et Geng) Zhu gen. et comb. nov. At the same time, the authors have confirmed that it is a kind of true fern. In association with the new taxon there are Sphenophyllum emarginatum, Bowmanites laxus, Lobatannularia lingulata, Chansitheca palaeosilvana, Ch. cf. kidstonii, Pecopteris anderssonii, P.echinata, P. ?taiyuanensis , P.unita, Danaeites mirabilis, Fascipteris densata, Cladophlebis ?permica , Compsopteris contracta, Gigantonoclea chingii, G.fukienensis, G.sp., ?G . sp., Gigantopteris cordata, G.nicotianaefolia, Taenioteris serrulata, T.szei, T.tingii, Radicites sp.1, R.sp. 2 etc. The present flora is more simplified than the others which discovered from different areas in the same geological time. Based on the grouping of these fossil plants, the geological era of the present flora must be late Early Permian to early Late Permian. Moreover it is emphasized that the gigantopterides is only but an artificial compound taxon not only including seed ferns but also true ferns and others.