短命植物舟果荠形态学的研究 Ⅱ繁殖器官的解剖构造

A Study of Morphology of Short-Lived Plant Tauscheria lasiocarpa Fisch. ex DC. Ⅱ. Anatomical Structure of the Reproductive Organ

本研究采用石蜡切片法,对舟果荠的繁殖器官解剖,观察到:萼片和药隔中都有大和较大的薄壁细胞,药壁4层,中层1层退化较早,绒毡层腺质型,消失晚,一朵花的花药间减数分裂多数不同步,四分孢子四面体形。成熟的花粉粒多数是3细胞2沟。4个花粉囊只有2个发育较好。倒生胚珠,双珠被,有明显的珠被绒毡最,薄珠心型,四分孢子直立式,反足细胞受精前退化,属蓼型胚囊,胚胎发育属十字花型。核型胚乳,无丝分裂。

The study of anatomical structure of reproductive organ Of Tauscheria lasiocarpa was observed by means of paraffin section: The sepal and connective all have larger and large parenchyma cells. The Wall of anther is 4-layered. A single middle layer degenerates early, The glandular tapetum disappears tardily. The meiosis is not completely synchronous between the anthers of a flower, Arrangement of microspores is tetrahedral. The most of mature pollens are 3 celled with 2 meridional valleculas exine. There are 4 pollen sacs in a anther but only two of them are fully developed. The ovule is anatropous, bitegmic and tenuinucellar and it has apparent integument tapetum. The tetrades of macrospores are erectstyle. The antipodal cells degenerate before fertilization. The development of embryosac belongs to the Polygonum type. Embryogeny follows the Cruciferae type and endosperm the nuclear type. The nucleus of endosperm is amitosis and division is not synchronous.