黄花菜大孢子和小孢子的发生及雌、雄配子体的发育

Megasporogenesis,Microsporogenesis and Development of Female and Male Gametophyte of Hemerocallis citrine Baroni

用常规石蜡切片技术,对黄花菜大孢子和小孢子的发生及雌、雄配子体发育进行了研究,结果表明:花药壁发育为单子叶型。小孢子母细胞减数分裂的胞质分裂为连续型。小孢子四分体排列为左右对称型和四面体型。成熟花粉为二细胞型。花粉为椭球形,具有一个萌发沟。生殖细胞呈现前期至早中期核形态。双珠被、薄珠心、倒生胚珠。珠心表皮下的雌性孢原直接发育为大孢子母细胞。减数分裂产生线形四分体。合点端大孢子为功能大孢子。雌配子体发育为蓼型。记录了雌、雄配子体发育的对应关系。成熟胚囊里,反足细胞早期退化,助细胞无钩状结构并具发达的丝状器以及中央细胞内没有明显大液泡是黄花菜胚囊发育的重要特征。

This paper studies the megasporogenesis, microsporogenesis male megasporogenesis by way of paraltin wax of HemerocaUis citrine Baroni. and development of female and The results are as follows： The development of anther wall follows Monocotyledonous type. Successive cytokinesis in microspore mother cell accompanies meiosis to produce tetrahedral tetrad as well grains are 2-celled when shed, with one germinal furrow. as medianly zygomorphic tetrad. Ellipsoidal pollen At this time, the nucleus in generative cell present the state of prophase or metaphase of mitosis. Ovule is bitegminous, temuinucellatae and anatropous. The archesporial cell below nucellus epiderm functions directly as the megaspore mother cell. The chalazal megaspore of linear tetrad develops into a Polygonum type embryo sac. Recorded relative development stage of pistil and stamen. In mature embryo sac, antipodal cell degenerates early. The main characteristics of the development of embryo sac in this species are that synergids without crook are abounded in filiform apparatus and there is no eminent vacuole in central cell.