摘要
研究了在中国最为广泛栽培的毛竹(Phyllostachys edulis(Cart.)H.de Lehaie)的地下茎的创伤反应。创伤后的第1天,在伤口附近没有明显反应;2d之后,创口附近可以观察到生理反应的代谢物,在后生木质部的导管中以及在基本组织细胞的胞间隙出现粘状物质,这些粘状物质具有果胶特性;创伤后1周,创口附近的筛管及基本组织中的短细胞的细胞壁变成木质化。同时,基本组织中的长细胞的内壁出现新的次生壁的沉积;2周后,创伤反应的组织与未受创伤的组织之间的区别变得更加明显;4周后,一些导管完全充满了粘性物质,但没有观察到侵填体。由于细胞壁的木质化及酚类物质的填充,筛管完全失去了功能。对创伤后6周的材料进行观察的结果表明,其创伤反应的范围不再扩展,在创伤组织与未受伤组织之间的基本组织的细胞壁变得相当厚,从而在两者之间形成一道屏障。毛竹地下茎的这种创伤反应与毛竹竹竿的创伤反应基本上是一致的,只是略有不同。
The wound reactions in the rhizome of Phyllostachys edulis (Carr.) H. de Lehaie, the main cultivated bamboo species in China, were studied. Three days after wounding metabolites of physiological reactions were observed near the wound. In the metaxylem vessels slimy substances appeared . After one week the walls of sieve tubes and short parenchyma cells near the wound edge became lignified. Additional lamellae were deposited on the inner wall of the longer parenchyma cells. Two weeks after wounding a more clear distinction was visualized between the wound modificated tissue and the undamaged one. After four weeks some vessels were filled up with the slimy material. Tyloses were not observed, however. Sieve tubes became functionless due to encrustation with phenolic substances and lignification. These wound responses of the rhizome were basically the same as those of the bamboo culm only with some specific differences.
基金
The work has been supported partly by the Deutsche Forschungsgemeinschaft(DFG) and partly by the International Foundation of soience(IFS),Swoden.
关键词
毛竹
地下茎
创伤反应
Bamboo, Rhizome, Anatomy, Wound Reaction
