中国蹄盖蕨属的研究I─属下分类

A REVISION OF THE CHINESE ATHYRIUM ROTH (I) ─ SUBGENERIC CLASSIFICATION OF THE GENUS

蹄盖蕨属是蕨类植物中分类最困难的大属之一，中国、喜马拉雅和日本为其分布中心。隶属于本属的种实际上是由蹄盖蕨科或广义蹄盖蕨属中其他属被划分出去以后剩余下来的种类集合而成的“大杂烩”，因此，它们被放在一起并非是一个单系起源的自然类群；属下不同类群分化的程度相差也很大，有的类群进化活跃，近缘种很多，有的类群则很孤立，所以属下单位所包括的种数相差悬殊；孢子周壁表面褶皱的有无在本属的属下分类中占有重要的位置。已发表的中国蹄盖蕨属植物名称约有３００多个，它们初步被处理为１１７种和一些变种和杂种，本文建议把这些种分为１４个组和１５个系。

Athyrium is one of the largest, complex and extremely difficult genera in taxonomic treatment. The area of China, Himalaya and Japan is the geographic distribution center of the genus. In fact, the species in this genus sensu stricto (Ching) are the remainders after other genera have been separated from the family or the genus Athyrium sensu lato. They are somewhat like a 'mixed stew', but not a natural monophyletic group. Some of the subgeneric unites are very active in evolution and with large number of related species, while some others are isolated or unique. Therefore, the size of the infrageneric groups are not equal. Presence or absence of folds on perispore surface is an important character for the subgeneric classification. There are more than 300 Chinese Athyrium species name published, they have been preliminarily treated as 117 species with a number of varieties and hybrids. They are suggested to be classified into 14 sections and 15 series in present paper.