竹柏属和罗汉松属叶输导组织的观察

Conducting Tissue of Leaves in Nageia and Podocarpus

该研究对罗汉松属和竹柏属植物叶横切面和纵切面输导组织结构进行观察。结果表明:罗汉松属叶1条主脉,主脉维管束木质部管胞发达,转输组织为苏铁型,转输组织管胞等径,栅栏组织和海绵组织之间的副转输组织管胞发达;竹柏属叶多条平行叶脉,维管束木质部管胞相对不发达,转输组织为红豆杉型,转输组织管胞纵向伸长,栅栏组织和海绵组织之间不存在副转输组织。结合罗汉松属植物主脉维管束下面有3个树脂道,叶肉分化为栅栏组织和海绵组织,而竹柏属植物每个叶脉维管束下面有1个树脂道,叶肉分化不明显,两面都有栅栏组织,叶肉中还具有大量石细胞等特征,认为罗汉松属和竹柏属在叶脉、叶形态和结构方面存在明显差异,2个属分开是比较合理的。

The conducting tissue structure of transverse and longitudinal sections was observed on leaves of Podocarpus and Nageta. Results showed ： in Podocarpus leaves, there is only one midrib, the xylem tracheid of midrib vascular bundle is multi-form, transfusion tissue belongs to Cycas-type and transfusion tracheids are isodiametric, the accessory transfusion tracheids between palisade tissue and sponge tissue are developed ; in Nageia leaves, there are plenty of parallel leaves, the xylem tracheids of each vein are relatively simple, transfusion tissue belongs to Taxus-type and transfusion tracheids are longer in longitudinal section than that in transverse section, the accessory transfusion tissue between pal- isade tissue and sponge tissue is absent. Considering other differences that in leaves of Podocarpus there are three resin ducts under vascular bun- dle of midrib, mesophyll cells are differentiated into palisade tissue and sponge tissue; in leaves of Nageia, there is only one resin duct under vascular bundle in each vein and no obvious differentiation in mesophyll cells, palisade tissue can be found on both sides, and sclereids can also be found in mesophyll tissue. The anatomical differences of leaf veins and mesophylls between Nageia and Podocarpus mentioned above support the viewpoint that Nageia and Podocarpus are two independent genera.