摘要
为进一步探讨香料烟叶腺毛的形态发生和分泌活动规律,为优质香料烟的开发提供理论依据,试验通过石蜡切片法观察了腺毛的形态发生过程,利用光学显微镜和扫描电镜观察了腺毛的分泌活动。结果发现,长柄腺毛和短柄腺毛最早均产生于茎尖高约100～200μm的第1对叶原基上,腺毛原初细胞大部分发育成长柄腺毛;长柄腺毛和短柄腺毛由原表皮细胞突起分化产生,经过不同的形态发生过程逐渐形成。长柄腺毛一般在烟叶欠熟时分泌活动开始增强,至叶片适熟期进入分泌高峰,腺头细胞外出现大量胶状分泌物,但部分腺毛分泌活动滞后,在烟叶调制时进入分泌高峰;短柄腺毛从叶片生长成熟到调制陈化均没有观察到腺头细胞外有分泌物。
The morphology development and secretion activity of glandular trichomes in oriental tobacco were investigated by paraffin section, optical microscope and scanning electron microscope, respectively. The results showed that both primary long and short trichomes originated from the first and second leaf primordium at 100-200μm, while the initial triehome cells generally developed into long trichomes. The differentiation cells bulging from protoderm developed into long or short trichomes through different morphogenesis evolved. The secretion of long trichomes grandually strengthened in the ver-matured leaves, and came to climax with lots of gummy exudate overflowed from gland cells when the leaves were proper mature. However, the secretion of some trichomes delayed, entering the climax during curing of the leaves. The gummy exudate was hardly observed in the short trichomes during the growth, mature, curing and aging of the leaves.
出处
《中国烟草科学》
CSCD
北大核心
2010年第1期22-27,共6页
Chinese Tobacco Science
基金
国家烟草专卖局重点科研项目“优质香料烟生产关键技术研究”(110199901002)
关键词
香料烟
腺毛
形态发生
分泌活动
oriental tobacco
glandular trichome
morphology development
secretion activity
