子午岭桥山林区柴松群落主要种群种间联结性研究

Interspecific Association of Dominant Populations of Pinus tabulaeformis f.shekanensis Communities in Qiaoshan Forest Area

基于2×2联列表,应用χ2检验、W检验、联结系数AC和共同出现百分率PC等方法对陕西黄土高原子午岭桥山林区柴松群落乔木层14种主要种群、灌木层23种主要种群和草本层24个主要种群的种间联结显著性和关联强度进行了定量测定,以揭示柴松森林群落不同结构层次中主要植物种群的种间关系。结果显示:(1)柴松群落乔木层、灌木层和草本层主要物种的总体关联性均为正相关。(2)乔木层中柴松与辽东栎、山杏、山桃,辽东栎与山杏、山桃,白桦与山杨之间有明显的正联结;山杏与山桃之间有明显的负联结。(3)灌木层中灰栒子和五加、陕西荚蒾与野葡萄、陕西荚蒾与蒙古荚蒾、盘叶忍冬与忍冬以及草本层中凤毛菊与黄芩、铁杆蒿与地榆、悬钩子与薯蓣、荩草与油芒、菝葜与糙苏、防风与黄花蒿之间存在较为明显的正联结。(4)根据种间联结特征,将14个主要乔木树种划分为4个生态种组,其中柴松和辽东栎为第1生态种组,山楂、白桦、白蜡和山杨为第2生态种组,山桃、茶条槭、毛梾和侧柏为第3生态种组,杜梨、山杏、山荆子和盐肤木为第4生态种组。

Based on the 2×2 table,series of approaches including variance analysis,χ2-test,percentage co-occurrence（PC） and association coefficient（AC） were used to analyze the interspecific associations of dominant tree species,shrub populations and herb populations of Pinus tabulaeformis f.shekanensis communities in Qiaoshan Forest Region.The results show that interspecific association community in general for arbor layer,shrub layer and herb layer in P.tabulaeformis f.shekanensis communities had positive association.In the arbor layer,P.tabulaeformis f.shekanensis was siginificantly positive associated with Quercus wutaishanica,Prunus armeniaca and P.davidiana;Q.wutaishanica with Prunus armeniaca and Prunus davidiana.However,Betula platyphylla was siginificantly negative associated with Populus davidiana.The inter-specific association between a couple of species was siginificantly positive,such as Cotoneaster acutifolius and Acanthopanax gracilistylus,Viburnum schensianum and Vitis amurensis or V.mongolicum,Lonicera tragophylla and L.japonica in shrub layer.Meanwhile,Saussurea japonica and Scutellaria baicalensis,Artemisia gmelinii and Sanguisorba officinalis,Rubus corchorifolius and Dioscorea matsudai,Arthraxon hispidus and Eccoilopus cotulifer,Smilax china and Phlomis umbrosa,Fructus lyeii and Artemisia annua in herb layer showed the same results.Based on the interspecific association,14 dominant species could be divided into four ecogroups of species.The first group was Pinus tabulaeformis f.shekanensis and Quercus wutaishanica;The second group was Crataegus pinnatifida,Betula platyphylla,Fraxinus velutina and Populus davidiana;The third group was Prunus davidiana,Acer ginnala,Cornus walteri and Platycladus orientalis;The last was Pyrus betulaefolia,Prunus armeniaca,M.baccata and Rhus chinensis.