大豆籽粒形成期^(14)C同化物的分配和源库调节效应的研究

Partitioning of ^(14)C-Assimilates and Effects of Source-Sink Manipulation at Seed-Filling in Soybean

在籽粒形成期,大豆不同叶位(14)~C-同化物的运转和分配具有明显的“局部供应性”,供应中心是本叶腋内荚粒.然而,植株不同节位叶片的(14)~C-同化物在各器官的分配又具一定差异,倒13叶分配到茎和荚粒的(14)~C-同化物高于倒1、4、7叶.叶片(14)~C-同化物向其他各节位不同器官分配的量较少,并存在明显的“同侧运输性”,主要分配到同侧节位荚粒,而且,向下运输量高于向上运输量.下部叶的(14)~C-同化物向相邻节各器官分配的范围和数量最广和最大,中部叶次之,顶部叶最窄和最小.摘除部分叶和荚后,保留叶和去荚叶的(14)~C同化物向本叶腋内荚粒分配的量减少,向相邻节位各器官分配的(14)C同化物增加,特别是向同侧节位荚粒分配的(14)~C同化物增加显著,而且,向下运转和分配的增加量高于向上增加量.

At pod-filling of soybean, translocation and partitioning of 14C-assimilates in leaves at different positions showed apparent local supplying. Leaf 14C-assimilates were mainly translocated into the seed in its axil. However, different translocation and partitioning of 14C-assimilates existed among the leaves at different positions. The 13th leaf (from upper to lower) partitioned more 14C-assimilates in its stem and seed than the 1st, 4th and 7th leaf did. Leaf partitioned little 14C-assimilates in organs at other nodal point and mainly in same lateral seed near it. However, 14C-assimilates were translocated more into lower organs than into higher organs. Markedly different range and amount of 14C-assimilates translocated into other organs occurred among leaves at different nodal points. The lower leaves had maximum range and amount of 14C-assimilates translocated into other nodal points, the middle leaves second, upper leaves third and top leaves least. Reducing partial leaves and pods decreased the partition of 14C-assimilates in retaining leaves and leaves without pods in seed of its axil and increased that in organs of same nodal point near this leaf. Also, 14C-assimilates partitioned in lower organs increased more notably than that in higher organs.