论单基因的分离负荷

On Segregational Load

本文引进两个配子选择系数、三个基因互作系数及互作跃变函数描述分离负荷，统一了近四十年来的理论纷争，得到如下结果：(1）只有当对两个配子的选择系数相等时，种群才能平衡．对平衡种群，以往关于分离负荷的结论是成立的，而且种群的适宜度保持不变；(2）当两配子选择系数不等时，种群不可能平衡，此时选择下分离负荷从Ｆ１代开始逐代减小，种群向杂合子纯合方向进化，从而解释了许多过去不能解释的现象．

This paper introduces two gametic selection coefficients and three gene interac- tion coefficients and a jump-function of gene interaction into theory described segregational load, and unites confused wrangling in theory lasted nearly forty years, and obtains such results as below: 1) For an equilibrium population the past conclusions about segregational load are no fault with a few exceptions, the equilibrium is probable only when S1=S2 and the fitness of population is unchangable. 2) If S1≠S2. the equilibrium is unprobable. For nonequilibrium population ever since F1-generation the segregational load decreases from one generation to next generation under the selection, the population is able to rise in power once again for short time. This way. many phe nomenon what was no able to explain in past are explained.