革苞菊花蜜腺的发育解剖学研究

Developmental and Anatomic Studies on the Floral Nectaries in Tugarinovia mongolica

革苞菊 (Tugarinoviamongolica)花蜜腺为筒状 ,围绕花柱基部着生 ,属于花柱蜜腺类型。蜜腺中无维管束分布 ,只具分泌表皮和泌蜜组织。分泌表皮细胞排列整齐 ,表皮细胞之间分散分布着许多气孔 ,气孔的孔下室发达。蜜腺基部细胞与花柱维管束相邻。随着蜜腺的发育 ,泌蜜组织的细胞结构和储藏物质呈规律性变化。原蜜汁的来源主要是由花柱基部维管束提供多糖 ,经过泌蜜组织加工成蜜汁后 ,由分泌表皮上的气孔将蜜汁排出体外。

Tugarinovia mongolica has a tubular stylar nectary located at the base of the style.The nectary, which lacks special vascular bundleis,is composed of secretory epidermis and nectariferous tissue. Stomata are present in the epidermis and there are well developed space under them.The basal cells of the nectary are adjacent to the vascular bundle of the style. During the development of nectary, both the structure and content of cell change regularly. The pre nectar is provided by the vascular bundle at the base of the style. Nectar is formed in nectariferous tissue,and it is excreted from the stoma. The relationship between the structure of the nectary and evolution is discussed.