间伐对日本落叶松人工林林下更新木本植物组成和生长影响的时间效应

Temporal effects of thinning on the composition and growth of regenerated woody plants in Larix kaempferi plantations

明确间伐对针叶人工林林下更新阔叶木本植物组成和生长影响的时间效应,可为促进人工纯林向针阔混交林转化、进而解决纯林生产与生态功能无法兼顾的问题提供参考。本研究以我国北方重要针叶人工林——日本落叶松人工林为例,比较间伐后短期(1~3年)、中期(4~9年)和长期(>9年)人工林林下更新木本植物组成的差异;并选择具有较高重要值且与落叶松互惠共生的3种更新树种蒙古栎、色木槭、糠椴,分析不同间伐时期人工林内光照(郁闭度)与更新木本植物生长的关系。结果表明:在间伐样地内更新的阔叶木本植物达46种,不同间伐时期样地内共有且占据优势的乔木树种为色木槭,灌木树种为忍冬和卫矛。随间伐后时间的推移,更新物种数逐渐减少,但乔灌比呈增大的趋势;中性树种占据优势地位。间伐对更新树种生长影响的时间效应受树种耐荫性的影响,糠椴的基径和树高均高于蒙古栎和色木槭。随间伐后时间的延长,糠椴的基径、色木槭的树高对光照响应较敏感,分别倾向于采用"耐荫策略"和"避荫策略"以适应间伐后变化的环境。间伐对日本落叶松人工林林下更新阔叶木本植物组成和生长的影响具有明显的时间效应;在制定以促进人工林林下树种更新为目标的间伐措施时,应该考虑适当延长间伐间隔,从而保证糠椴和色木槭等阔叶树幼苗的生长,使其进入林冠层,促进形成针阔混交林,实现人工林的可持续发展。

Understanding the temporal effects of thinning on the composition and growth of regene-rated broadleaved woody species in coniferous plantations can provide profound references for promoting the conversion of monoculture plantations into mixed conifer-broadleaved forests,which could solve the problem that the production and ecological functions of monoculture plantations cannot be balanced.We compared the composition of regenerated woody plant species in Larix kaempferi plantation with short-term(1-3 years),medium-term(4-9 years)and long-term(>9 years)after thinning.Furthermore,we selected three regenerated tree species with higher importance value and reciprocal symbiosis with L.kaempferi,which differed in shade tolerance,including shade-intolerant species Quercus mongolica,intermediate shade-tolerant species Acer mono,and shade-tolerant species Tilia mandschurica.We analyzed the relationships between light conditions(i.e.,canopy density)and the growth(i.e.,base diameter and height)of those species in L.kaempferi plantation with different terms after thinning.The results showed that 46 species of regene-rated broadleaved woody plants were recorded in thinned plantations.The common and dominant tree in different terms after thinning was A.mono,and the shrub species were Lonicera japonica and Euonymus alatus.With the increasing time after thinning,species richness of regenerated trees decreased,but the ratio of tree to shrub species increased and the intermediate shade-tolerant tree species took the dominant position.The temporal effect of thinning on the growth of three tree species was affected by shade tolerance ability.Basal diameter and height of T.mandschurica were higher than those of Q.mongolica and A.mono.With the increases of time after thinning,basal diameter of T.mandschurica and height of A.mono were more sensitive to light,indicating that there were respectively"a shade tolerance strategy"and"a shade avoidance strategy"to adapt to the post-thinning environment.The effects of thinning on the composition and growth of regenerated broadleaved woody species in L.kaempferi plantations were significantly time-sensitive.When deve-loping thinning measures to promote the regeneration of broadleaved trees in plantations,we should consider to extend the thinning interval appropriately to ensure the growth of broadleaved tree seedlings(e.g.,T.mandschurica and A.mono)and accelerate their migration into the canopy layer.This would promote the formation of mixed conifer-broadleaved forests and eventually realize the sustainable development of plantations.