The diurnal temperature range(DTR) serves as a vital indicator reflecting both natural climate variability and anthropogenic climate change. This study investigates the historical and projected multitemporal DTR varia...The diurnal temperature range(DTR) serves as a vital indicator reflecting both natural climate variability and anthropogenic climate change. This study investigates the historical and projected multitemporal DTR variations over the Tibetan Plateau. It assesses 23 climate models from phase 6 of the Coupled Model Intercomparison Project(CMIP6) using CN05.1 observational data as validation, evaluating their ability to simulate DTR over the Tibetan Plateau. Then, the evolution of DTR over the Tibetan Plateau under different shared socioeconomic pathway(SSP) scenarios for the near,middle, and long term of future projection are analyzed using 11 selected robustly performing models. Key findings reveal:(1) Among the models examined, BCC-CSM2-MR, EC-Earth3, EC-Earth3-CC, EC-Earth3-Veg, EC-Earth3-Veg-LR,FGOALS-g3, FIO-ESM-2-0, GFDL-ESM4, MPI-ESM1-2-HR, MPI-ESM1-2-LR, and INM-CM5-0 exhibit superior integrated simulation capability for capturing the spatiotemporal variability of DTR over the Tibetan Plateau.(2) Projection indicates a slightly increasing trend in DTR on the Tibetan Plateau in the SSP1-2.6 scenario, and decreasing trends in the SSP2-4.5, SSP3-7.0, and SPP5-8.5 scenarios. In certain areas, such as the southeastern edge of the Tibetan Plateau, western hinterland of the Tibetan Plateau, southern Kunlun, and the Qaidam basins, the changes in DTR are relatively large.(3) Notably, the warming rate of maximum temperature under SSP2-4.5, SSP3-7.0, and SPP5-8.5 is slower compared to that of minimum temperature, and it emerges as the primary contributor to the projected decrease in DTR over the Tibetan Plateau in the future.展开更多
In a convective scheme featuring a discretized cloud size density, the assumed lateral mixing rate is inversely proportional to the exponential coefficient of plume size. This follows a typical assumption of-1, but it...In a convective scheme featuring a discretized cloud size density, the assumed lateral mixing rate is inversely proportional to the exponential coefficient of plume size. This follows a typical assumption of-1, but it has unveiled inherent uncertainties, especially for deep layer clouds. Addressing this knowledge gap, we conducted comprehensive large eddy simulations and comparative analyses focused on terrestrial regions. Our investigation revealed that cloud formation adheres to the tenets of Bernoulli trials, illustrating power-law scaling that remains consistent regardless of the inherent deep layer cloud attributes existing between cloud size and the number of clouds. This scaling paradigm encompasses liquid, ice, and mixed phases in deep layer clouds. The exponent characterizing the interplay between cloud scale and number in the deep layer cloud, specifically for liquid, ice, or mixed-phase clouds, resembles that of shallow convection,but converges closely to zero. This convergence signifies a propensity for diminished cloud numbers and sizes within deep layer clouds. Notably, the infusion of abundant moisture and the release of latent heat by condensation within the lower atmospheric strata make substantial contributions. However, this role in ice phase formation is limited. The emergence of liquid and ice phases in deep layer clouds is facilitated by the latent heat and influenced by the wind shear inherent in the middle levels. These interrelationships hold potential applications in formulating parameterizations and post-processing model outcomes.展开更多
Alpine wetland is one of the typical underlying surfaces on the Qinghai–Tibet Plateau.It plays a crucial role in runoff regulation.Investigations on the mechanisms of water and heat exchanges are necessary to underst...Alpine wetland is one of the typical underlying surfaces on the Qinghai–Tibet Plateau.It plays a crucial role in runoff regulation.Investigations on the mechanisms of water and heat exchanges are necessary to understand the land surface processes over the alpine wetland.This study explores the characteristics of hydro-meteorological factors with in situ observations and uses the Community Land Model 5 to identify the main factors controlling water and heat exchanges.Latent heat flux and thermal roughness length were found to be greater in the warm season(June–August)than in the cold season(December–February),with a frozen depth of 20–40 cm over the alpine wetland.The transfers of heat fluxes were mainly controlled by longwave radiation and air temperature and affected by root distribution.Air pressure and stomatal conductance were also important to latent heat flux,and soil solid water content was important to sensible heat flux.Soil temperature was dominated by longwave radiation and air temperature,with crucial surface parameters of initial soil liquid water content and total water content.The atmospheric control factors transitioned to precipitation and air temperature for soil moisture,especially at the shallow layer(5 cm).Meanwhile,the more influential surface parameters were root distribution and stomatal conductance in the warm season and initial soil liquid water content and total water content in the cold season.This work contributes to the research on the land surface processes over the alpine wetland and is helpful to wetland protection.展开更多
To reveal the characteristics of evapotranspiration and environmental control factors of typical underlying surfaces(alpine wetland and alpine meadow)on the Qinghai-Tibetan Plateau,a comprehensive study was performed ...To reveal the characteristics of evapotranspiration and environmental control factors of typical underlying surfaces(alpine wetland and alpine meadow)on the Qinghai-Tibetan Plateau,a comprehensive study was performed via in situ observations and remote sensing data in the growing season and non-growing season.Evapotranspiration was positively correlated with precipitation,the decoupling coefficient,and the enhanced vegetation index,but was energy-limited and mainly controlled by the vapor pressure deficit and solar radiation at an annual scale and growing season scale,respectively.Compared with the non-growing season,monthly evapotranspiration,equilibrium evaporation,and decoupling coefficient were greater in the growing season due to lower vegetation resistance and considerable precipitation.However,these factors were restricted in the alpine meadow.The decoupling factor was more sensitive to changes of conductance in the alpine wetland.This study is of great significance for understanding hydro-meteorological processes on the Qinghai-Tibetan Plateau.展开更多
There are six micronuclear divisions during conjugation of Paramecium caudatum:three prezygotic and three postzygotic divisions.Four haploid nuclei are formed during the first two meiotic prezygotic divisions.Usually ...There are six micronuclear divisions during conjugation of Paramecium caudatum:three prezygotic and three postzygotic divisions.Four haploid nuclei are formed during the first two meiotic prezygotic divisions.Usually only one meiotic product is located in the paroral cone(PC)region at the completion of meiosis,which survives and divides mitotically to complete the third prezygotic division to yield a stationary and a migratory pronucleus.The remaining three located outside of the PC degenerate.The migratory pronuclei are then exchanged between two conjugants and fuse with the stationary pronuclei to form synkarya,which undergo three successive divisions(postzygotic divisions).However,little is known about the surviving mechanism of the PC nuclei.In the current study,stage-specific appearance of cytoplasmic microtubules(cMTs)was indicated during the third prezygotic division by immunofluorescence labeling with anti-alpha tubulin antibodies surrounding the surviving nuclei,including the PC nuclei and the two types of prospective pronuclei.This suggested that cMTs were involved in the formation of a physical barrier,whose function may relate to sequestering and protecting the surviving nuclei from the major cytoplasm,where degeneration of extra-meiotic products occurs,another important nuclear event during the third prezygotic division.展开更多
There are six micronuclear divisions during conjugation of Paramecium caudatum: three prezygotic and three postzygotic divisions. Four haploid nuclei are formed during the first two meiotic prezygotic divisions. Usua...There are six micronuclear divisions during conjugation of Paramecium caudatum: three prezygotic and three postzygotic divisions. Four haploid nuclei are formed during the first two meiotic prezygotic divisions. Usually only one meiotic product is located in the paroral cone (PC) region at the completion of meiosis, which survives and divides mitotically to complete the third prezygotic division to yield a stationary and a migratory pronucleus. The remaining three located outside of the PC degenerate. The migratory pronuclei are then exchanged between two conjugants and fuse with the stationary pronuclei to form synkarya, which undergo three successive divisions (postzygotic divisions). However, little is known about the surviving mechanism of the PC nuclei. In the current study, stage-specific appearance of cytoplasmic microtubules (cMTs) was indicated during the third prezygotic division by immunofluorescence labeling with anti-alpha tubulin antibodies surrounding the surviving nuclei, including the PC nuclei and the two types of prospective pronuclei. This suggested that cMTs were involved in the formation of a physical barrier, whose function may relate to sequestering and protecting the surviving nuclei from the major cytoplasm, where degeneration of extra-meiotic products occurs, another important nuclear event during the third prezygotic division.展开更多
Using the cloud radar,ground observations,and ECMWF Reanalysis v5(ERA5)data,we investigate the factors influencing nighttime precipitation during summer in the Yushu area of the Tibetan Plateau(TP).The cloud top heigh...Using the cloud radar,ground observations,and ECMWF Reanalysis v5(ERA5)data,we investigate the factors influencing nighttime precipitation during summer in the Yushu area of the Tibetan Plateau(TP).The cloud top height(CTH),cloud base height(CBH),and liquid water content(LWC)are compared between non-precipitation and precipitation days.The results show that the average CTH on precipitation days in Yushu is below 10 km above ground level(AGL)in the daytime,whereas it exceeds 10 km AGL at night,with the maximum at 2300 BT(Beijing Time).The CBH is in-phase with the dewpoint spread.The precipitation intensity and CTH are in-phase with the LWC.The hourly averaged precipitation intensity and convective available potential energy in ERA5 reach their maximums at2100 BT,which is 3 h ahead of their observed counterparts.There is descending motion in the mid day on non-precipitation days,whereas there is ascending motion at night on precipitation days.In addition,the horizontal wind direction in the lower level(below 5000 m)shows clockwise rotation from morning to night.Wind shear occurs in the mid level of the atmosphere,accompanied by a subtropical westerly jet in the upper level.The difference in horizontal wind speed between 200 and 500 hPa is positively related to the LWC,thereby contributing to the formation of upper-level cloud.展开更多
The three-river source region plays an important role on China’s ecological security and Asia’s water supply. Historically, the region has experienced severe ecological degradation due to climate change and human ac...The three-river source region plays an important role on China’s ecological security and Asia’s water supply. Historically, the region has experienced severe ecological degradation due to climate change and human activities. Reasonable simulations of the energy and water cycles are essential to predict the responses of land surface processes to future climate change. Current land surface models involve empirical functions that are associated with many parameters. These parameter uncertainties will largely affect the simulation when applied to a new domain. The Community Land Model(CLM) is a widely used land surface model, and version 5.0 is the newest version. Compared to the prior version CLM4.5, CLM5.0 has largely updated plant hydraulic and stomatal conductance schemes. How these changes affect parameter sensitivities is unknown. In our work, we tested 17 key parameters in CLM4.5 and 19 parameters in CLM5.0 at two eddy flux sites in the three-river source region: the Maqu and Maduo sites. We adopted the simplest one-at-a-time changes on each parameter and quantified their sensitivities by the parameter effect(PE).We found that the Maqu site was more sensitive to vegetation parameters, while the Maduo site was more sensitive to the initial soil water content in both CLM4.5 and CLM5.0. This is because Maduo grid cell has wetland that does not respond to vegetation parameters in CLM, which may not reflect the reality. Further model development on wetland vegetation parameterization is important. Our validation on the default simulation showed CLM5.0 did not always improve the simulations. The largest difference between CLM5.0 and CLM4.5 was that soil moisture(SM) showed a much stronger decrease in response to a higher leaf area index(LAI) in CLM5.0 than in CLM4.5, suggesting that SM is more sensitive to vegetation changes in CLM5.0.展开更多
基金supported by The Second Tibetan Plateau Scientific Expedition and Research (STEP) program(Grant No. 2019QZKK0102)the National Natural Science Foundation of China (Grant No. 41975135)+1 种基金the Natural Science Foundation of Sichuan,China (Grant No. 2022NSFSC1092)funded by the China Scholarship Council。
文摘The diurnal temperature range(DTR) serves as a vital indicator reflecting both natural climate variability and anthropogenic climate change. This study investigates the historical and projected multitemporal DTR variations over the Tibetan Plateau. It assesses 23 climate models from phase 6 of the Coupled Model Intercomparison Project(CMIP6) using CN05.1 observational data as validation, evaluating their ability to simulate DTR over the Tibetan Plateau. Then, the evolution of DTR over the Tibetan Plateau under different shared socioeconomic pathway(SSP) scenarios for the near,middle, and long term of future projection are analyzed using 11 selected robustly performing models. Key findings reveal:(1) Among the models examined, BCC-CSM2-MR, EC-Earth3, EC-Earth3-CC, EC-Earth3-Veg, EC-Earth3-Veg-LR,FGOALS-g3, FIO-ESM-2-0, GFDL-ESM4, MPI-ESM1-2-HR, MPI-ESM1-2-LR, and INM-CM5-0 exhibit superior integrated simulation capability for capturing the spatiotemporal variability of DTR over the Tibetan Plateau.(2) Projection indicates a slightly increasing trend in DTR on the Tibetan Plateau in the SSP1-2.6 scenario, and decreasing trends in the SSP2-4.5, SSP3-7.0, and SPP5-8.5 scenarios. In certain areas, such as the southeastern edge of the Tibetan Plateau, western hinterland of the Tibetan Plateau, southern Kunlun, and the Qaidam basins, the changes in DTR are relatively large.(3) Notably, the warming rate of maximum temperature under SSP2-4.5, SSP3-7.0, and SPP5-8.5 is slower compared to that of minimum temperature, and it emerges as the primary contributor to the projected decrease in DTR over the Tibetan Plateau in the future.
基金supported by the Second Tibetan Plateau Scientific Expedition and Research Program (STEP) (Grant No.2019QZKK010203)the National Natural Science Foundation of China (Grant No.42175174 and 41975130)+1 种基金the Natural Science Foundation of Sichuan Province (Grant No.2022NSFSC1092)the Sichuan Provincial Innovation Training Program for College Students (Grant No.S202210621009)。
文摘In a convective scheme featuring a discretized cloud size density, the assumed lateral mixing rate is inversely proportional to the exponential coefficient of plume size. This follows a typical assumption of-1, but it has unveiled inherent uncertainties, especially for deep layer clouds. Addressing this knowledge gap, we conducted comprehensive large eddy simulations and comparative analyses focused on terrestrial regions. Our investigation revealed that cloud formation adheres to the tenets of Bernoulli trials, illustrating power-law scaling that remains consistent regardless of the inherent deep layer cloud attributes existing between cloud size and the number of clouds. This scaling paradigm encompasses liquid, ice, and mixed phases in deep layer clouds. The exponent characterizing the interplay between cloud scale and number in the deep layer cloud, specifically for liquid, ice, or mixed-phase clouds, resembles that of shallow convection,but converges closely to zero. This convergence signifies a propensity for diminished cloud numbers and sizes within deep layer clouds. Notably, the infusion of abundant moisture and the release of latent heat by condensation within the lower atmospheric strata make substantial contributions. However, this role in ice phase formation is limited. The emergence of liquid and ice phases in deep layer clouds is facilitated by the latent heat and influenced by the wind shear inherent in the middle levels. These interrelationships hold potential applications in formulating parameterizations and post-processing model outcomes.
基金supported by the National Natural Science Foundation of China(Grant Nos.42005075,41975130)Natural Science Foundation of Gansu Province(Grant No.21JR7RA047)+1 种基金Open Research Fund Program of Plateau Atmosphere and Environment Key Laboratory of Sichuan Province(Grant No.PAEKL-2022-K03)the State Key Laboratory of Cryospheric Science(Grant No.SKLCS-ZZ-2023,SKLCS-ZZ-2022).
文摘Alpine wetland is one of the typical underlying surfaces on the Qinghai–Tibet Plateau.It plays a crucial role in runoff regulation.Investigations on the mechanisms of water and heat exchanges are necessary to understand the land surface processes over the alpine wetland.This study explores the characteristics of hydro-meteorological factors with in situ observations and uses the Community Land Model 5 to identify the main factors controlling water and heat exchanges.Latent heat flux and thermal roughness length were found to be greater in the warm season(June–August)than in the cold season(December–February),with a frozen depth of 20–40 cm over the alpine wetland.The transfers of heat fluxes were mainly controlled by longwave radiation and air temperature and affected by root distribution.Air pressure and stomatal conductance were also important to latent heat flux,and soil solid water content was important to sensible heat flux.Soil temperature was dominated by longwave radiation and air temperature,with crucial surface parameters of initial soil liquid water content and total water content.The atmospheric control factors transitioned to precipitation and air temperature for soil moisture,especially at the shallow layer(5 cm).Meanwhile,the more influential surface parameters were root distribution and stomatal conductance in the warm season and initial soil liquid water content and total water content in the cold season.This work contributes to the research on the land surface processes over the alpine wetland and is helpful to wetland protection.
基金This work is financially supported by the National Natural Science Foundation of China(Grant Nos.42005075 and 41530529),the Second Tibetan Plateau Scientific Expedition and Research Program(STEP)(No.2019QZKK0605)the State Key Laboratory of Cryospheric Science(Grant Nos.SKLCS-ZZ-2020 and SKLCS-ZZ-2021)+1 种基金Foundation for Excellent Youth Scholars of"Northwest Institute of Eco-Environment and Resources",CAS(Grant No.FEYS2019020)The authors declare no competing interest in this paper.Our cordial gratitude should be extended to anonymous reviewers and the Editors for their professional and pertinent comments on this manuscript.
文摘To reveal the characteristics of evapotranspiration and environmental control factors of typical underlying surfaces(alpine wetland and alpine meadow)on the Qinghai-Tibetan Plateau,a comprehensive study was performed via in situ observations and remote sensing data in the growing season and non-growing season.Evapotranspiration was positively correlated with precipitation,the decoupling coefficient,and the enhanced vegetation index,but was energy-limited and mainly controlled by the vapor pressure deficit and solar radiation at an annual scale and growing season scale,respectively.Compared with the non-growing season,monthly evapotranspiration,equilibrium evaporation,and decoupling coefficient were greater in the growing season due to lower vegetation resistance and considerable precipitation.However,these factors were restricted in the alpine meadow.The decoupling factor was more sensitive to changes of conductance in the alpine wetland.This study is of great significance for understanding hydro-meteorological processes on the Qinghai-Tibetan Plateau.
基金National Natural Science Foundation of China(31071881)Research Development Foundation of Zhejiang A&F University(2009FK67).
文摘There are six micronuclear divisions during conjugation of Paramecium caudatum:three prezygotic and three postzygotic divisions.Four haploid nuclei are formed during the first two meiotic prezygotic divisions.Usually only one meiotic product is located in the paroral cone(PC)region at the completion of meiosis,which survives and divides mitotically to complete the third prezygotic division to yield a stationary and a migratory pronucleus.The remaining three located outside of the PC degenerate.The migratory pronuclei are then exchanged between two conjugants and fuse with the stationary pronuclei to form synkarya,which undergo three successive divisions(postzygotic divisions).However,little is known about the surviving mechanism of the PC nuclei.In the current study,stage-specific appearance of cytoplasmic microtubules(cMTs)was indicated during the third prezygotic division by immunofluorescence labeling with anti-alpha tubulin antibodies surrounding the surviving nuclei,including the PC nuclei and the two types of prospective pronuclei.This suggested that cMTs were involved in the formation of a physical barrier,whose function may relate to sequestering and protecting the surviving nuclei from the major cytoplasm,where degeneration of extra-meiotic products occurs,another important nuclear event during the third prezygotic division.
基金Foundation items: National Natural Science Foundation of China (31071881) Research Development Foundation of Zhejiang A & F University (2009FK67).
文摘There are six micronuclear divisions during conjugation of Paramecium caudatum: three prezygotic and three postzygotic divisions. Four haploid nuclei are formed during the first two meiotic prezygotic divisions. Usually only one meiotic product is located in the paroral cone (PC) region at the completion of meiosis, which survives and divides mitotically to complete the third prezygotic division to yield a stationary and a migratory pronucleus. The remaining three located outside of the PC degenerate. The migratory pronuclei are then exchanged between two conjugants and fuse with the stationary pronuclei to form synkarya, which undergo three successive divisions (postzygotic divisions). However, little is known about the surviving mechanism of the PC nuclei. In the current study, stage-specific appearance of cytoplasmic microtubules (cMTs) was indicated during the third prezygotic division by immunofluorescence labeling with anti-alpha tubulin antibodies surrounding the surviving nuclei, including the PC nuclei and the two types of prospective pronuclei. This suggested that cMTs were involved in the formation of a physical barrier, whose function may relate to sequestering and protecting the surviving nuclei from the major cytoplasm, where degeneration of extra-meiotic products occurs, another important nuclear event during the third prezygotic division.
基金Supported by the Second Tibetan Plateau Scientific Expedition and Research Program(2019QZKK0102)National Natural Science Foundation of China(41975130 and 42005072)+1 种基金National Key Research and Development Program of China(2018YFC1505702)Sichuan Provincial Innovation Training Program for College Student(S202010621048 and S202110621004)。
文摘Using the cloud radar,ground observations,and ECMWF Reanalysis v5(ERA5)data,we investigate the factors influencing nighttime precipitation during summer in the Yushu area of the Tibetan Plateau(TP).The cloud top height(CTH),cloud base height(CBH),and liquid water content(LWC)are compared between non-precipitation and precipitation days.The results show that the average CTH on precipitation days in Yushu is below 10 km above ground level(AGL)in the daytime,whereas it exceeds 10 km AGL at night,with the maximum at 2300 BT(Beijing Time).The CBH is in-phase with the dewpoint spread.The precipitation intensity and CTH are in-phase with the LWC.The hourly averaged precipitation intensity and convective available potential energy in ERA5 reach their maximums at2100 BT,which is 3 h ahead of their observed counterparts.There is descending motion in the mid day on non-precipitation days,whereas there is ascending motion at night on precipitation days.In addition,the horizontal wind direction in the lower level(below 5000 m)shows clockwise rotation from morning to night.Wind shear occurs in the mid level of the atmosphere,accompanied by a subtropical westerly jet in the upper level.The difference in horizontal wind speed between 200 and 500 hPa is positively related to the LWC,thereby contributing to the formation of upper-level cloud.
基金Supported by the Strategic Priority Research Program of Chinese Academy of Sciences(XDA20050102)National Natural Science Foundation of China(41975135 and 41975130)。
文摘The three-river source region plays an important role on China’s ecological security and Asia’s water supply. Historically, the region has experienced severe ecological degradation due to climate change and human activities. Reasonable simulations of the energy and water cycles are essential to predict the responses of land surface processes to future climate change. Current land surface models involve empirical functions that are associated with many parameters. These parameter uncertainties will largely affect the simulation when applied to a new domain. The Community Land Model(CLM) is a widely used land surface model, and version 5.0 is the newest version. Compared to the prior version CLM4.5, CLM5.0 has largely updated plant hydraulic and stomatal conductance schemes. How these changes affect parameter sensitivities is unknown. In our work, we tested 17 key parameters in CLM4.5 and 19 parameters in CLM5.0 at two eddy flux sites in the three-river source region: the Maqu and Maduo sites. We adopted the simplest one-at-a-time changes on each parameter and quantified their sensitivities by the parameter effect(PE).We found that the Maqu site was more sensitive to vegetation parameters, while the Maduo site was more sensitive to the initial soil water content in both CLM4.5 and CLM5.0. This is because Maduo grid cell has wetland that does not respond to vegetation parameters in CLM, which may not reflect the reality. Further model development on wetland vegetation parameterization is important. Our validation on the default simulation showed CLM5.0 did not always improve the simulations. The largest difference between CLM5.0 and CLM4.5 was that soil moisture(SM) showed a much stronger decrease in response to a higher leaf area index(LAI) in CLM5.0 than in CLM4.5, suggesting that SM is more sensitive to vegetation changes in CLM5.0.