Remote sensing of quality traits in cereal and arable production systems:A review

Cereal is an essential source of calories and protein for the global population.Accurately predicting cereal quality before harvest is highly desirable in order to optimise management for farmers,grading harvest and categorised storage for enterprises,future trading prices,and policy planning.The use of remote sensing data with extensive spatial coverage demonstrates some potential in predicting crop quality traits.Many studies have also proposed models and methods for predicting such traits based on multiplatform remote sensing data.In this paper,the key quality traits that are of interest to producers and consumers are introduced.The literature related to grain quality prediction was analyzed in detail,and a review was conducted on remote sensing platforms,commonly used methods,potential gaps,and future trends in crop quality prediction.This review recommends new research directions that go beyond the traditional methods and discusses grain quality retrieval and the associated challenges from the perspective of remote sensing data.

Genome-wide association study exposed the pleiotropic genes for yield-related and oil quality traits in Brassica napus L.

Oilseed rape(Brassica napus L.) is an allotetraploid(AACC,2n=38) crop,valued for its edible oil and protein content.seed yield and nutritional composition of rapeseed are influenced by its yield and oil quality traits.However,the genetic basis of yield-related and oil-quality traits remain ambiguous.A panel of 266 diversified oilseed rape accessions was genotyped using 223 simple sequence repeat(SSR) markers covering all 19 chromosomes to identify significant markers associated with yield and quality traits.Twelve yield-related and six quality traits were investigated in two consecutive years(2014 and 2015),with three replications in two environments(Changshun,CS;and Qinghe,QH).Using the model GLM with population structure and kinship(Q+K),a total of 25 significant SSR markers(P <0.001) were detected to be associated with these twelve yield-related and six quality traits,explaining 4.56%-19.17% of the phenotypic variation for each trait.Based on these markers,BnaA03g23490D, BnaC09g46370D,BnaA07g37150D,BnaA01g32590D, and BnaC09g37280D were identified as pleiotropic genes controlling multiple traits.These candidate genes illustrated the potential for the genetic understanding of yield and oil quality traits.Most importantly,these significant markers can be used for marker-assisted breeding of oilseed rape in different environments.

Analysis of Correlation between Yield and Quality Traits of Quality Hybrid Rapeseed

[Objective] This study aimed to investigate the correlation between yield and quality traits of rape. [Method] Taking ＂Sanbei 98＂ and ＂Youyan 599＂ as ex- periment materials, correlation between yield and quality traits of Brassica napus L. cultivars was explored by orthogonal rotation method. [Result] The results showed that： 1 In both the two varieties-Sanbei98 and Youyan599, erucic acid content had the largest coefficient of variation, but its average content was lower than 2% in both the two varieties （0.90% in Sanbei98 and 1.24% in Youyan599）, and the palmitic acid content had the smallest coefficient of variation. 2 In terms of the dif- ferences in quality traits at different yield levels, only the mean of eicosenoic acid content revealed significant difference in Sanbei98, and only the mean of seed-pro- tein content revealed significant difference in Youyan599, and there was no signifi- cant difference in any combined quality traits in both the two varieties. 3 The yield was negatively correlated with the erucic acid content or oil content, and posi- tively correlated with the content of seed-protein, linoleic acid, or linolenic acid, and the coefficient of correlation between the yield and either of the contents of glucosi- nolates, oleic acid, palmitic acid, stearic acid, and eicosenoic acid was smaller. ~, when the yield of a variety increased considerably, there was a certain decreasing trend in erucic acid content, oil content, the total amount of erucic chain fatty acids （El ＆ Er）, while there was an increasing trend in seed-protein content, linoleic acid content, linolenic acid content, saturated fatty acids （SFA）, the total amount of fat and protein （P ＆ F）, unsaturation index （IUFA）, unsaturated fatty acids （UFA）, the total amount of 18 carbon fatty acids （18-C FA）, unsaturated fatty acids [UFA/（Ei ＆ Er） ], and the sum of oleic acid and linoleic acid （OI ＆ Li）. 5 When the seed-pro- tein content is increased, the quality of rape oil cake is also improved; when the unsaturated index is increased, the oil stability is decreased. The increase in the total amount of unsaturated fatty acids, oleic acid and linoleic acid helps to increase in the total amount of fatty acids absorbed by human bodies. [Conclusion] This study provides a theoretical foundation for the cultivation of high-yielding and top- quality hybrid rape cultivars.

Research on the Agronomic,Quality Traits and the Corresponding Background Genes Distribution of Wheat Germplasm Ningchun 4 and Its Parents

[Objective] The aim of this study was to observe the agronomic,yield and quality traits of Ningchun 4 and its parents under the same environmental conditions,as well as to carry out molecular detection on the background genes distribution of the corresponding traits.[Method]Ningchun 4 and its parents Sonora 64,Hongtu,Abbondanza,Quality were used as materials to detect the agronomic and quality traits,as well as to analyze the genetic variation laws by molecular determination method.[Result]Ningchun 4 had inherited the advantages of bigger spike,red and hard grain from Sonora 64 and higher 1 000-grain weight from Hongtu.However,it had also inherited the disadvantages of late-maturing from Sonora 64 and lower tillering ability from Hongtu;the grain quality of Ningchun 4 was slightly lower than the Sonora 64;Ningchun 4 had high quality subunit of 5＋10,which had good dough rheological properties.[Conclusion]Ningchun 4 had inherited the long photoperiod characteristics and no-resistant to slow-leaf rusting and stripe rust characteristics from Hongtu and low PPO activity,high yellow pigments content from parents.

Correlation between Carotenoids Content and Several Quality Traits of Wheat Varieties

A total of 31 wheat varieties（lines） released in different years in Huanghuai Area were selected as materials so as to study the carotenoids content,lipoxygenase activity, and some other quality traits of wholemeal. Results showed that carotenoid content and other quality traits had highly significant differences among varieties. Carotenoids content was significantly positive correlated with watersoluble pentosan and yellowness. Carotenoids content was significantly negative correlated with peak viscosity, hold through, breakdown, final viscosity, unit weight,formation time, and was negative correlated with peak time, protein content and stability time. According to cluster analysis, carotenoid contents in 31 varieties were divided into three major groups： one group had common carotenoid content, which included 19 varieties such as Meng 0318; one group had relatively low carotenoid content, which included 10 varieties such as Luomai23; and the last group had relatively high carotenoid content, which included 2 varieties such as Huaimai22. The average carotenoid contents of the three groups were 1.93, 1.07 and 2.99 mg/kg respectively.

Fatty Acid Composition and Seed Quality Traits of the Transgenic Rapeseed W-4(Brassica napus L.) with Down-regulated Expression of fad2 Gene

To clarify the effect of down-regulated expression of fad2 gene on the seed nutritional quality of rapeseed, the fatty acid composition, amino acid composi- tion, oil content, protein content, crude fiber content and glucosinolate content in the seeds of both transgenic line W-4 and its control Westar were compared. The re- sults showed that the oleic acid content in W-4 was 86.03%±0.20%, which was 29.36% higher than that in the control （P〈0.01）; the linoleic acid content was 2.86%± 0.01%, which was reduced by 84.03% compared with that in the control （P〈0.01）; the linolenic acid content in W-4 was 3.04%±0.04%, reduced by 57.60% （P〈0.01）; the palmitic acid content in W-4 was 3.23%±0.07%, reduced by 18.63% （P〈0.01）; the eicosenoic acid content in W-4 was increased by 18.46% compared with that in the control （P〈0.01）; the erucic acid content in W-4 was increased by 13.15% （P〈 0.05）; there was no significant difference in stearic acid content between the treat- ment and control groups （P〉0.05）. The amino acid composition analysis showed that total 18 amino acids, including 8 essential amino acids, were detected in both W-4 and Westar; there were no significant differences in contents of the 18 amino acids between the treatment and control groups except that of tyrosine （P〉0.05）; the contents of oil, proteins, glucosinolates and crude fiber in W-4 were 45.40%± 0.17% （P〉0.05）, 18.18%±0.91% （P〉0.05）, 18.20%±1.21% （P〉0.05） and 12.29%± 0.04% （P〉0.05）, respectively. All the results above showed that the down-regulated expression of fad2 had great effects on fatty acid composition and accumulation in rapeseed seeds, but had no significant effects on other seed quality traits, such as oil content, protein content, crude fiber content and glucosinolate content.

Effects of Soil and Environment on Quality Traits of Purple-fleshed Sweetpotato

For the study of main traits of purple-flashed sweetpotato in different soil and ecological environments, 18 cultivars of purple-flashed sweetpotato with different enthocyenin contents were planted at Yancheng and Xuzhou separately. The main quality traits were determined. The results showed the quality traits of purple-fleshed sweetpotato were significantly influenced by soil and ecological environments. Geno- type （G）, environment （E） and genotype x environment （GxE） interaction had signif- icantiy effects on the contents of anthocyanin, dry matter, starch, protein, reducing sugar and soluble sugar in storage roots. Anthocyanin content was most easily af- fected by the environment. The E effect on anthocyanin content was greater than G and GxE effects. However, the GxE interaction effects on other 5 quality traits were greater than G and E effects. The 6 quality traits were significantly different at 1% level under different environments respectively. The contents of dry matter, starch and anthocyenin were higher in Yancheng than in Xuzhou, while the contents of protein, reducing sugar and soluble sugar were higher in Xuzhou. The variation co- efficients of all quality traits at Yancheng and Xuzhou were in the same order： an- sugar〉protein〉starch, the variation coefficient of anthocyanin content was higher in Xuzhou, and the variation coefficients of other 5 traits were higher in Yancheng. Correlation analysis showed that the reducing sugar and solu- ble sugar contents were negatively correlated with starch content, and had no sig- nifiant correlation with anthocyanin and protein contents. Dry matter content and starch content were positively correlated, and anthocyanin content and other quality traits had no significant correlation.

Effect of HMW-GS 6 + 8 and 1.5 + 10 from Synthetic Hexaploid Wheat on Wheat Quality Traits

To determine the effect of 6 ＋ 8 and 1.5 ＋ 10 HMW-GS of synthetic hexaploid wheat （SHW） on main quality parameters of wheat, a set of recombinant inbred lines （RILs） derived from the cross between a SHW with N, 6 ＋ 8, 1.5 ＋ 10 HMW-GS and a cultivar Chuanyu 12-1 （CY 12-1） with 1, 7 ＋ 8, 2 ＋ 12 were planted in three environments in 2005 and 2006 and totally 16 quality parameters were tested for each line. Significant differences in all tested quality parameters but flour yield were observed between the two parents. The mean values of the RILs were intermediate to the parents for grain and protein parameters and some farinograph parameters, flour water absorption （FWA）, and farinograph softening （SOF） but beyond parents at dough stability time （DST）, breakdown time （BRT）, quality number （QN）, noodle score （NS）, and loaf volume （LOV）. All of the quality traits, especially in grain hardness （GH）, zeleny sedimentation volume （SED）, and most of farinograph parameters had significant difference between the different HMW-GS components. The effects of different alleles of HMW-GS at same locus （Glu-A1 or Glu-B1 or GIu-D1） on the different quality parameters were also different and affected by the other two loci. For most of parameters tested, 6 ＋ 8 was better than 7 ＋ 8 and there was no difference between 1.5 ＋ 10 and 2 ＋ 12. End-use quality was greatly influenced by components of HMW-GS. The components of 1, 6 ＋ 8, 1.5 ＋ 10 had the highest LOV and bread score （BS） values, whereas the components of 1, 7＋ 8 and 1.5 ＋ 10 had the highest NS values. Noodle score performed a positive linear relationship with falling number （FN） and its relationships to other quality parameters were affected by environments. Loaf volume had a significant negative relationship to SOF and positive associations with most of quality parameters. It could be concluded that HMW-GS 6＋ 8 from SHW had better overall quality characteristics than 7 ＋ 8, whereas the effects of 1.5 ＋ 10 on quality was different in respect to quality parameters and the HMW-GS components. Synthetic hexaploid wheat with subunits 6 ＋ 8 and 1.5 ＋ 10 had the potentials to improve the end-use quality of wheat cultivars.

Identification of QTLs Conferring Agronomic and Quality Traits in Hexaploid Wheat

The availability of elite germplasm resources with high yield and quality potentials is very important for development of cultivars in wheat. Thus, seeking such resources has been the continuous effort of breeder community. In this study, genetic analysis of a novel resource, Triticum spelta line CSCR6, from Australia was made by use of a recombination inbred line （RIL） population of 82 individuals from the cross between CSCR6 and another Australian hexaploid wheat cultivar, Lang. Data of a multiple environmental test was employed to genetically dissect quantitative trait loci （QTL） for agronomic traits such as plant height （PH）, spike length （SL）, spikelet per spike （SPI）, grain number per spike （GNS） and thousand grains weight （TGW） and for quality traits including grain protein content （GPC）, gluten content （GC）, grain hardness （GH）, falling number （FN） and sedimentation value （SV）. A 24 QTLs with additive effects were detected for all the investigated traits, and were located on chromosomes 1B, 1D, 2B, 3A, 3B, 3D, 4B, 5A, 5B, 7A, and 7B, respectively. Some QTLs located on 2B and 4B showed higher explanation of phenotypic variances and were not obviously interacted with environment. A QTL in the marker interval of wPT-5334-wPT-4918 （near the locus barc 0199） on 4B gave the highest contribution ratio of 30.76% on PH, while Qgpc-4B and Qgc-4B gave 13.07 and 14.70% contribution ratio on GPC and GC, respectively. Qph-2B, Qgns-2B, and Qgpc-2B showed 13.36, 10.00, and 10.79% contribution ratio on PH, GNS and GPC, respectively. Also, a QTL on 5A, Qsl- 5A, could explain 25.12% of phenotypic variance on SL. For most of agronomic and quality traits, CSCR6 alleles produced increase effects. The fact that a number of loci affecting the investigated traits were detected in T. spelta line CSCR6 revealed that it could offer a new opportunity for the manipulation of these traits in wheat breeding programs.

The influence of drought stress on malt quality traits of the wild and cultivated barleys

As a major abiotic stress,drought causes instability and deterioration of malt barley quality.There is distinct difference among barley cultivars in the responses of the main malt quality traits to drought stress.In the previous study,we identified some Tibetan wild barley accessions with relatively less change of malt quality traits under drought.In this study,we examined the impact of drought stress during grain filling stage on grain weight and several important malt quality traits,including total protein content,β-glucan content,limit dextrinase activity,β-amylase activity,and protein fractions in four barley genotypes(two Tibetan wild accessions and two cultivars).Drought treatment reduced grain weight,β-glucan content,and increased total protein content,β-amylase activity.These changes differed among barley genotypes and treatments,and are closely associated with grain filling process and kernel weight.All the results indicated Tibetan wild barley had great potential for developing drought tolerant barley cultivars.Relatively stable kernel weight or filling process under water stress should be highlighted in malt barley breeding in order to reduce the effect of water stress on malt barley quality.

QTL and genetic analysis controlling fiber quality traits using paternal backcross population in upland cotton

Background:Genetic improvement in fiber quality is one of the main challenges for cotton breeders.Quantitative trait loci(QTL)mapping provides a powerful approach to dissect the molecular mechanism in fiber quality traits.In present study,F14 recombinant inbred line(RIL)population was backcrossed to paternal parent for a paternal backcross(BC/P)population,deriving from one upland cotton hybrid.Three repetitive BC/P field trials and one maternal backcross(BC/M)field trial were performed including both two BC populations and the original RIL population.Results:In total,24 novel QTLs are detected for fiber quality traits and among which 13 QTLs validated previous results.Thirty-five QTLs in BC/P populations explain 5.01%–22.09%of phenotype variation(PV).Among the 35 QTLs,23 QTLs are detected in BC/P population alone.Present study provides novel alleles of male parent for fiber quality traits with positive genetic effects.Particularly,qFS-Chr3–1 explains 22.09%of PV in BC/P population,which increaseds 0.48 cN·tex−1 for fiber strength.A total of 7,2,8,2 and 6 QTLs explain over 10.00%of PV for fiber length,fiber uniformity,fiber strength,fiber elongation and fiber micronaire,respectively.In RIL population,six common QTLs are detected in more than one environment:qFL-Chr1–2,qFS-Chr5–1,qFS-Chr9–1,qFS-Chr21–1,qFM-Chr9–1 and qFM-Chr9–2.Two common QTLs of qFE-Chr2–2(TMB2386-SWU12343)and qFM-Chr9–1(NAU2873-CGR6771)explain 22.42%and 21.91%of PV.The region between NAU4034 and TMB1296 harbor 30 genes(379 kb)in A05 and 42 genes(49 kb)in D05 for fiber length along the QTL qFL-Chr5–1 in BC/P population,respectively.In addition,a total of 142 and 46 epistatic QTLs and QTL×environments(E-QTLs and QQEs)are identified in recombinant inbred lines in paternal backcross(RIL-P)and paternal backcross(BC/P)populations,respectively.Conclusions:The present studies provide informative basis for improving cotton fiber quality in different populations.

QTL Mapping for Fiber Quality Traits Based on a Dense Genetic Linkage Map with SSR,TRAP,SRAP and AFLP Markers in Cultivated Tetraploid Cotton

Cotton is one of the most important economic crops in the world,and it provides natural fiber for the textile industry.With the advancement of the textile technology and increased consumption demands on cotton fiber,both cotton yield and quality should be enhanced.However,cotton yield

Identification of candidate genes controlling fiber quality traits in upland cotton through integration of meta-QTL,significant SNP and transcriptomic data

Background:Meta-analysis of quantitative trait locus(QTL)is a computational technique to identify consensus QTL and refine QTL positions on the consensus map from multiple mapping studies.The combination of meta-QTL intervals,significant SNPs and transcriptome analysis has been widely used to identify candidate genes in various plants.Results:In our study,884 QTLs associated with cotton fiber quality traits from 12 studies were used for meta-QTL analysis based on reference genome TM-1,as a result,74 meta-QTLs were identified,including 19 meta-QTLs for fiber length;18 meta-QTLs for fiber strength;11 meta-QTLs for fiber uniformity;11 meta-QTLs for fiber elongation;and 15 meta-QTLs for micronaire.Combined with 8589 significant single nucleotide polymorphisms associated with fiber quality traits collected from 15 studies,297 candidate genes were identified in the meta-QTL intervals,20 of which showed high expression levels specifically in the developing fibers.According to the function annotations,some of the 20 key candidate genes are associated with the fiber development.Conclusions:This study provides not only stable QTLs used for marker-assisted selection,but also candidate genes to uncover the molecular mechanisms for cotton fiber development.

Analysis of genetic effects and correlations for nutrient quality traits of indica rice in different environments

Understanding the variation for the expressionof genes in different environments is one of themajor goals in qualitative genetics. In this pa-per, the genetic models for quantitative traitsof endosperm in cereal crops were used to eval-uate the seed, cytoplasm and maternal geneticeffects as well as the genotype × environment(GE) interaction effects, and to predict thebreeding value of parents and genotypic corre-lation for nutrient quality traits of indica rice (Oryza sativa L.).

Molecular Markers in Improvement of Fiber Quality Traits in Cotton

Cotton is the worlds leading natural fiber crop,and it is the cornerstone of textile industries worldwide.The cotton industry is confronted with problems in cost of production and

Analysis of genetic relationships between grain milling quality traits and plant agronomic traits of rice

Since seed and maternal plant are in different generations,the grain milling traits and plant agronomic traits of rice are based on differentgenetic models.Using the genetic modelswhich could estimate covariance componentsbetween two traits with unequal design matri-ces,we analyzed the gentic relationships be-

Genetic analysis of quality traits in intersubspecies crosses of rice

Indica and japonica rices are two subspecies ofAsia cultivated rice.Their grain qualities exista significant difference.The objective of thispaper is to study the genetic control of qualitytraits in intersubspecies crosses of rice.In 1991,4 japonica varieties with widecompatibility,PC311,Zhenxiqiuguang,Lun-hui 422,and Ce 03 were crossed with 5 indicavarieties,3037,Zaoxiandang 18,IR36,Minghui 63,and Yangdao 4 according to NC(?)design.The 20 hybrids using japonica as ma-

Effect of Cold-Water Irrigation on Grain Quality Traits in japonica Rice Varieties from Yunnan Province,China

The response of grain quality traits to cold-water irrigation and its correlation with cold tolerance were studied in 11 japonica rice varieties from Yunnan Province, China. The results indicated that the response of grain quality traits to the cold-water stress varied with rice varieties and grain quality traits. Under the cold-water stress, grain width, chalky rice rate, whiteness, 1000-grain weight, brown rice rate, taste meter value, peak viscosity, trough viscosity, breakdown viscosity and final viscosity significantly decreased, whereas grain length-width ratio, head rice rate, alkali digestion value, protein content and setback viscosity markedly increased. However, the other traits such as grain length, amylose content, milled rice rate, peak viscosity time and pasting temperature were not significantly affected by the cold-water stress. Significant correlations were discovered between phenotypic acceptability and cold response indices of taste meter value, protein content, peak viscosity and breakdown viscosity. Therefore, it would be very important to improve the cold tolerance of Yunnan rice varieties in order to stabilize and improve their eating quality.

QTL Mapping for Quality Traits of Chinese Dry Noodle

The QTLs for quality traits of Chinese dry noodle （CDN） were mapped based on a population of recombinant inbred lines （RILs） derived from the cross between two Chinese winter wheat varieties, Chuan 35050 and Shannong 483. Sensory quality traits of CDN were tested according to the standard of Ministry of Commerce of P.R.China （SB/T 10137-93）, and the textural property traits were detected using a texture analyser （TA.XTplus）. We have obtained 8 putative QTLs for 6 sensory quality traits and 2 QTLs for textural property of CDN with a single QTL explainning 4.07-75.67% of phenotypic variations on chromosomes 1A, 1 D, 3D, 4A, and 6D. A cluster of 3 QTLs for palate, elasticity and smoothness of CDN was found near the Glu-D I locus on chromosome 1D with high contributions, The increasing effect come from Chuan 35050, and the relationship between the QTLs were positive. On chromosome 4A, co-location QTLs for stickiness and total score were detected in the region of Xwmc420-Xswes620-Xswes615. Their contributions were high and the increasing effects come from Shannong 483. A taste QTL QStas.sdau-4A.1 was obtained in Xsrap18-Xswes624c-Xissr25b-Xissr23b on chromosome 4A with the highest contribution of 75.67%. This QTL was a major gene and the increasing effects come from Shannong 483.

QTL Analysis for Seven Quality Traits of RIL Population in Japonica Rice Based on Three Genetic Statistical Models

QTL mapping for seven quality traits was conducted by using 254 recombinant inbred lines （RIL） derived from a japonica-japonica rice cross of Xiushui 79/C Bao. The seven traits investigated were grain length （GL）, grain length to width ratio （LWR）, chalk grain rate （CGR）, chalkiness degree （CD）, gelatinization temperature （GT）, amylose content （AC） and gel consistency （GC） of head rice. Three mapping methods employed were composite interval mapping in QTLMapper 2.0 software based on mixed linear model （MCIM）, inclusive composite interval mapping in QTL IciMapping 3.0 software based on stepwise regression linear model （ICIM） and multiple interval mapping with regression forward selection in Windows QTL Cartographer 2.5 based on multiple regression analysis （MIMR）. Results showed that five QTLs with additive effect （A-QTLs） were detected by all the three methods simultaneously, two by two methods simultaneously, and 23 by only one method. Five A-QTLs were detected by MCIM, nine by ICIM and 28 by MIMR. The contribution rates of single A-QTL ranged from 0.89% to 38.07%. All the QTLs with epistatic effect （E-QTLs） detected by MIMR were not detected by the other two methods. Fourteen pairs of E-QTLs were detected by both MCIM and ICIM, and 142 pairs of E-QTLs were detected by only one method. Twenty-five pairs of E-QTLs were detected by MCIM, 141 pairs by ICIM and four pairs by MIMR. The contribution rates of single pair of E-QTL were from 2.60% to 23.78%. In the Xiu-Bao RIL population, epistatic effect played a major role in the variation of GL and CD, and additive effect was the dominant in the variation of LWR, while both epistatic effect and additive effect had equal importance in the variation of CGR, AC, GT and GC. QTLs detected by two or more methods simultaneously were highly reliable, and could be applied to improve the quality traits in japonica hybrid rice.