A new genus and species of basal neoceratopsian dinosaur, Auroraceratops rugosus, is reported based on material from the Early Cretaceous Xinminpu Group in the Gongpoquan Basin of Gansu Province, China. Auroraceratops...A new genus and species of basal neoceratopsian dinosaur, Auroraceratops rugosus, is reported based on material from the Early Cretaceous Xinminpu Group in the Gongpoquan Basin of Gansu Province, China. Auroraceratops is represented by a nearly complete skull and low jaws, and different greatly from all other neoceratopsians by its considerable breadth of the nasals, fungi form expansion of the dorsal end of the lacrimal, highly developed rugosity of the jugal, dentary and surangular, and inflated, striated premaxillary teeth. The finding of Auroraceratops adds diversity and helps elucidate the evolution of basal neoceratopsian dinosaurs.展开更多
We herein describe an associated partial postcranial skeleton of a theropod dinosaur discovered in the Lower Cretaceous Xinminpu Group of the Yujingzi Basin, in the Jiuquan area of Gausu Province, northwestern China. ...We herein describe an associated partial postcranial skeleton of a theropod dinosaur discovered in the Lower Cretaceous Xinminpu Group of the Yujingzi Basin, in the Jiuquan area of Gausu Province, northwestern China. Features of its humerus, such as strongly expanded proximal and distal ends, a well developed medial tuberosity, distal condyles expressed on the humeral cranial surface, and a hypertrophied entepicondyle, definitively establish the therizinosauroid affinities of the specimen. It differs from other therizinosauroids in having a shallow, poorly demarcated glenoid fossa with a prominent rounded and striated tumescence on the dorsomedial surface of its scapular portion, and a pubis with a strongly concave cranial margin. It represents a new taxon, Suzhousaurus megatherioides gen. et sp. nov. Cladistic analysis recovers Suzhousaurus as the sister taxon of Nothronychus mckinleyi from the mid-Cretaceous of western North America; together, they are basal members of the Therizinosauroidea, more derived than the Early Cretaceous Falcarius and Beipiaosaurus but less derived than Alxasaurus and the Therizinosauridae. Along with "Nanshiungosaurus" bohlini from possibly coeval beds in the Mazongshan area of northern-most Gansu, Suzhousaurus represents one of the largest-known Early Cretaceous therizinosauroids, demonstrating that this clade attained considerable body size early in its evolutionary history.展开更多
Although the fossil record of non-avian dinosaurs from the Cretaceous of Antarctica is the poorest of any continent,fossils representing at least five major taxonomic groups(Ankylosauria,early-diverging Ornithopoda,Ha...Although the fossil record of non-avian dinosaurs from the Cretaceous of Antarctica is the poorest of any continent,fossils representing at least five major taxonomic groups(Ankylosauria,early-diverging Ornithopoda,Hadrosauridae,Titanosauria,and Theropoda)have been recovered.All come from Upper Cretaceous(Coniacian–Maastrichtian)marine and nearshore deposits belonging to the Gustav and Marambio groups of the James Ross Basin at the northern tip of the Antarctic Peninsula.The majority of these finds have come from the Campanian–Maastrichtian Snow Hill Island and López de Bertodano formations of James Ross and Vega islands.Given the rarity of Antarctic Cretaceous non-avian dinosaurs,discoveries of any fossils of these archosaurs,no matter how meager,are of significance.Here we describe fragmentary new ornithischian(ankylosaur and ornithopod)material from the upper Campanian–lower Maastrichtian Cape Lamb Member of the Snow Hill Island Formation and the Maastrichtian Sandwich Bluff Member of the López de Bertodano Formation.One of these specimens is considered to probably pertain to the holotypic individual of the early-diverging ornithopod Morrosaurus antarcticus.We also provide an up-to-date synthesis of the Late Cretaceous non-avian dinosaur record of the James Ross Basin and analyze the biostratigraphic occurrences of the various finds,demonstrating that most(including all named taxa and all reasonably complete skeletons discovered to date)occur within a relatively condensed temporal interval of the late Campanian to early Maastrichtian.Most or all James Ross Basin dinosaurs share close affinities with penecontemporaneous taxa from Patagonia,indicating that at least some continental vertebrates could disperse between southern South America and Antarctica during the final stages of the Mesozoic.展开更多
Recent studies demonstrate that many avialan features evolved incrementally prior to the origin of the group,but the presence of some of these features,such as bird-like brooding behaviours,remains contentious in non-...Recent studies demonstrate that many avialan features evolved incrementally prior to the origin of the group,but the presence of some of these features,such as bird-like brooding behaviours,remains contentious in non-avialan dinosaurs.Here we report the first non-avialan dinosaur fossil known to preserve an adult skeleton atop an egg clutch that contains embryonic remains.The preserved positional relationship of the adult to the clutch,coupled with the advanced growth stages of the embryos and their high estimated incubation temperatures,provides strong support for the brooding hypothesis.Furthermore,embryos in the clutch are at different developmental stages,suggesting the presence of asynchronous hatching—a derived feature even among crown-group birds—in non-avialan theropods.These findings demonstrate that the evolution of reproductive biology along bird-line archosaurs was a complex rather than a linear and incremental process,and suggest that some aspects of non-avialan theropod reproduction were unique to these dinosaurs.展开更多
Paleogene strata in the Huheboerhe area,Erlian Basin,Inner Mongolia,China have been subdivided into three lithological units:the Nomogen Formation,the Arshanto Formation,and the Irdin Manha Formation.At least 12 mamma...Paleogene strata in the Huheboerhe area,Erlian Basin,Inner Mongolia,China have been subdivided into three lithological units:the Nomogen Formation,the Arshanto Formation,and the Irdin Manha Formation.At least 12 mammal-bearing horizons have been recognized in these formations,of which 4 in the Nomogen Formation,6 in the Arshanto Formation,and 2 in the Irdin Manha Formation.Recent investigation proved that the "Houldjin Formation" recognized in this area by the Central Asiatic Expeditions(CAE) of the American Museum Natural History in the 1920s is actually the Irdin Manha Formation,while the "Irdin Manha Formation" of CAE is the Arshanto Formation.A recent paleomagnetic study suggests that the upper part of the Nomogen Formation is early Eocene in age and the Arshanto Formation is mainly early Eocene rather than middle Eocene as previously thought.The Gashatan,Bumbanian,and Arshantan land mammal ages are correlated respectively to the Thanetian,the early Ypresian,and the middle Ypresian through earliest Lutetian of the Geological Time Scale.These land mammal ages are also correlated with the late Tiffanian through Clarkforkian,the early Wasatchian,and the middle-late Wasatchian and most of the Bridgerian of the North American Land Mammal Ages.During the early Paleogene,the mammalian history of the Erlian Basin was dominated by the appearance of new mammalian families and the replacement of a variety of genera and species,corresponding to the gradual climatic changes during this time period.The abrupt emergence of several modern mammalian orders at the beginning of the Eocene is probably related to extreme climatic warming in relation to the Paleocene-Eocene Thermal Maximum.展开更多
文摘A new genus and species of basal neoceratopsian dinosaur, Auroraceratops rugosus, is reported based on material from the Early Cretaceous Xinminpu Group in the Gongpoquan Basin of Gansu Province, China. Auroraceratops is represented by a nearly complete skull and low jaws, and different greatly from all other neoceratopsians by its considerable breadth of the nasals, fungi form expansion of the dorsal end of the lacrimal, highly developed rugosity of the jugal, dentary and surangular, and inflated, striated premaxillary teeth. The finding of Auroraceratops adds diversity and helps elucidate the evolution of basal neoceratopsian dinosaurs.
基金We are grateful to Ji Qiang, Tang Zhilu, and Peter Dodson for their generous help. We also thank Zhang Yuqing for the preparation of FRDC-GSJB-99 Equal thanks go to every member of the field team. Early drafts of this paper benefited from discussions with Lindsay Zanno and Bucky Gates. Funding was provided by the Ministry of Science and Technology of China (973 Project: 2006CB701405) to Ji Qiang, the National Natural Science Foundation of China (40672007) the Hundred Talents Project of Ministry of Land and Resources of China to You Hailu, and Bureau of Geology and Mineral Resources of Gansu Province to Li Daqing.
文摘We herein describe an associated partial postcranial skeleton of a theropod dinosaur discovered in the Lower Cretaceous Xinminpu Group of the Yujingzi Basin, in the Jiuquan area of Gausu Province, northwestern China. Features of its humerus, such as strongly expanded proximal and distal ends, a well developed medial tuberosity, distal condyles expressed on the humeral cranial surface, and a hypertrophied entepicondyle, definitively establish the therizinosauroid affinities of the specimen. It differs from other therizinosauroids in having a shallow, poorly demarcated glenoid fossa with a prominent rounded and striated tumescence on the dorsomedial surface of its scapular portion, and a pubis with a strongly concave cranial margin. It represents a new taxon, Suzhousaurus megatherioides gen. et sp. nov. Cladistic analysis recovers Suzhousaurus as the sister taxon of Nothronychus mckinleyi from the mid-Cretaceous of western North America; together, they are basal members of the Therizinosauroidea, more derived than the Early Cretaceous Falcarius and Beipiaosaurus but less derived than Alxasaurus and the Therizinosauridae. Along with "Nanshiungosaurus" bohlini from possibly coeval beds in the Mazongshan area of northern-most Gansu, Suzhousaurus represents one of the largest-known Early Cretaceous therizinosauroids, demonstrating that this clade attained considerable body size early in its evolutionary history.
基金supported by NSF grants ANT-1142129 to MCL, ANT-1141820 to JAC2, ANT-1142104 to PMO, ANT- 0636639 and ANT-1142052 to R. MacPhee, and OPP-9615933 and ANT-0003844 to JAC1
文摘Although the fossil record of non-avian dinosaurs from the Cretaceous of Antarctica is the poorest of any continent,fossils representing at least five major taxonomic groups(Ankylosauria,early-diverging Ornithopoda,Hadrosauridae,Titanosauria,and Theropoda)have been recovered.All come from Upper Cretaceous(Coniacian–Maastrichtian)marine and nearshore deposits belonging to the Gustav and Marambio groups of the James Ross Basin at the northern tip of the Antarctic Peninsula.The majority of these finds have come from the Campanian–Maastrichtian Snow Hill Island and López de Bertodano formations of James Ross and Vega islands.Given the rarity of Antarctic Cretaceous non-avian dinosaurs,discoveries of any fossils of these archosaurs,no matter how meager,are of significance.Here we describe fragmentary new ornithischian(ankylosaur and ornithopod)material from the upper Campanian–lower Maastrichtian Cape Lamb Member of the Snow Hill Island Formation and the Maastrichtian Sandwich Bluff Member of the López de Bertodano Formation.One of these specimens is considered to probably pertain to the holotypic individual of the early-diverging ornithopod Morrosaurus antarcticus.We also provide an up-to-date synthesis of the Late Cretaceous non-avian dinosaur record of the James Ross Basin and analyze the biostratigraphic occurrences of the various finds,demonstrating that most(including all named taxa and all reasonably complete skeletons discovered to date)occur within a relatively condensed temporal interval of the late Campanian to early Maastrichtian.Most or all James Ross Basin dinosaurs share close affinities with penecontemporaneous taxa from Patagonia,indicating that at least some continental vertebrates could disperse between southern South America and Antarctica during the final stages of the Mesozoic.
基金supported by the Double First-Class Joint Program of Yunnan Science and Technology Department and Yunnan University (2018FY001-005)the China-Myanmar Joint Laboratory for Ecological and Environmental Conservation+2 种基金the University of Hong Kong Faculty of Science RAE Improvement Fundsupported by the CNRS Program INSU INTERRVIEthe National Natural Science Foundation of China(41688103)。
文摘Recent studies demonstrate that many avialan features evolved incrementally prior to the origin of the group,but the presence of some of these features,such as bird-like brooding behaviours,remains contentious in non-avialan dinosaurs.Here we report the first non-avialan dinosaur fossil known to preserve an adult skeleton atop an egg clutch that contains embryonic remains.The preserved positional relationship of the adult to the clutch,coupled with the advanced growth stages of the embryos and their high estimated incubation temperatures,provides strong support for the brooding hypothesis.Furthermore,embryos in the clutch are at different developmental stages,suggesting the presence of asynchronous hatching—a derived feature even among crown-group birds—in non-avialan theropods.These findings demonstrate that the evolution of reproductive biology along bird-line archosaurs was a complex rather than a linear and incremental process,and suggest that some aspects of non-avialan theropod reproduction were unique to these dinosaurs.
基金supported by National Basic Research Program of China (Grant No. 2006CB806400)National Natural Science Foundation of China (Grant No. 40532010)+1 种基金the Basic Work Program of MST of China (Grant No. 2006FY120300-15)Special Fund for Fossil Excavation and Preparation of Chinese Academy of Sciences,and US National Science Foundation (Grant Nos. EAR-0120727, BCS-0309800)
文摘Paleogene strata in the Huheboerhe area,Erlian Basin,Inner Mongolia,China have been subdivided into three lithological units:the Nomogen Formation,the Arshanto Formation,and the Irdin Manha Formation.At least 12 mammal-bearing horizons have been recognized in these formations,of which 4 in the Nomogen Formation,6 in the Arshanto Formation,and 2 in the Irdin Manha Formation.Recent investigation proved that the "Houldjin Formation" recognized in this area by the Central Asiatic Expeditions(CAE) of the American Museum Natural History in the 1920s is actually the Irdin Manha Formation,while the "Irdin Manha Formation" of CAE is the Arshanto Formation.A recent paleomagnetic study suggests that the upper part of the Nomogen Formation is early Eocene in age and the Arshanto Formation is mainly early Eocene rather than middle Eocene as previously thought.The Gashatan,Bumbanian,and Arshantan land mammal ages are correlated respectively to the Thanetian,the early Ypresian,and the middle Ypresian through earliest Lutetian of the Geological Time Scale.These land mammal ages are also correlated with the late Tiffanian through Clarkforkian,the early Wasatchian,and the middle-late Wasatchian and most of the Bridgerian of the North American Land Mammal Ages.During the early Paleogene,the mammalian history of the Erlian Basin was dominated by the appearance of new mammalian families and the replacement of a variety of genera and species,corresponding to the gradual climatic changes during this time period.The abrupt emergence of several modern mammalian orders at the beginning of the Eocene is probably related to extreme climatic warming in relation to the Paleocene-Eocene Thermal Maximum.