基金Key Program of the National Natural Science Fund(31130061)National Nature Science Foundation of China(31572278+7 种基金 3180198131470214)Central Finance Forestry Protection Grant Funds(2014-2016)Foundation of Shaanxi Academy of Science,China(2016K-202018nk-18)the Strategic Priority Research Program of the Chinese Academy of Sciences(XDB31020302)Key Research and Development Program of Shaanxi Province of China(2018PT-04 2018 ZDXM-NY-049)