基金supported by Pan-Third Pole Environment Study for a Green Silk Road of the Strategic Priority Research Program of Chinese Academy of Sciences(XDA20090000)the National Key Research and Development Program of China(2018YFA0606400)+1 种基金Part laboratory and fieldwork costs were supported by the National Natural Science Foundation of China(41772178,91747207,and 41620104007)Field expedition was supported by the Second Tibetan Plateau Scientific Expedition(2019QZKK0601)。