基金financially supported by the National Key R&D Program on Nano Science & Technology of the MOST (2017YFA0207301)the National Natural Science Foundation of China (21925110,21890751,91745113,12147105,22275205,and 22005284)+9 种基金the National Program for Support of Top-notch Young Professionals,the Fundamental Research Funds for the Central Universities (WK2060190084)the Youth Innovation Promotion Association CAS (2018500),the National Postdoctoral Program for Innovative Talents (BX20190307,BX20190308)the Major Program of Development Foundation of Hefei Centre for Physical Science and Technology (2016FXZY001)the Users with Excellence Project of Hefei Science Centre CAS (2018HSC-UE002)the CAS Project for Young Scientists in Basic Research (YSBR-070)the support from the beamline 1W1B of Beijing Synchrotron Radiation Facility (BSRF,Beijing,China)beamline BL12B-a of the National Synchrotron Radiation Laboratory (NSRL,Hefei,China)the Cryo-EM Centre at the University of Science and Technology of China for the EM facility supportpartially carried out at the USTC Centre for Micro and Nanoscale Research and Fabricationthe support from the Super Computer Centre of USTCSCC and SCCAS