基金GDAS' Project of Science and Technology Development(2019GDASYL-0103064)Project Funded by China Postdoctoral Science Foundation(2018M643032)+3 种基金Guangdong Basic and Applied Basic Research Foundation(2019A1515110795)Graduate and Postdoctoral Fund of Guangdong Institute of Applied Biological Resources(GIABR-pyjj201804)Science and Technology Planning Project of Guangdong(2017A020219002 and 2018B030324003)GDAS Special Project of Science and Technology Development(2018GDASCX-0107)。