基金supported by the Funds of the Chinese Academy of Sciences for Key Topics in Innovation Engineering(KZCX2-YW-216,KSCX2-YW-G-065,KSCX2-YW-G-073)Natural Science Funds of South China Sea Institute of Oceanology for Young Scholar(SQ200903)+1 种基金Funds of Frontier Areas of SCSIO (LYQY200805)Open Project Program of Key Laboratory of Marine Bio-resources Sustainable Utilization(LMB091013)