基金supported by the National Key R&D Program of China(2017YFA0505500)National Natural Science Foundation of China(11771152,12026608,11901203,31930022,and 31771476)+7 种基金Guangdong Basic and Applied Basic Research Foundation(2019B151502062,and 2021A1515012317)Strategic Priority Research Program of the Chinese Academy of Sciences(XDB38040400)Shanghai Municipal Science and Technology Major Project(2017SHZDZX01)Japan Society for the Promotion of Science KAKENHI(15H05707)Japan Science and Technology Agency Moonshot R&D(JPMJMS2021)Japan Agency for Medical Research and Development(JP20dm0307009)UTokyo Center for Integrative Science of Human Behavior(CiSHuB)the International Research Center for Neurointelligence(WPI-IRCN)at The University of Tokyo Institutes for Advanced Study(UTIAS).