基金This study was supported by the National Natural Science Foundation of China to Hao Yu(NSFC-32060113/31702006)the Natural Science Foundation of Guizhou Province to Hao Yu([2020J1Y081)+1 种基金Guizhou Education University Academic Discipline Project(2019YLPYXKB01)Guizhou provincial first-class major(biological science)project(Education department of guizhou province[2019]46)。