基金supported by the National Science Fund for Distinguished Young Scientists (No. 30925008)the Major International (Regional) Joint Research Project (No. 31010103901)+1 种基金the CAS-IOZ Innovation Program (KSCX2-EW-J-2) by a grant (No. O529YX5105) from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences to F.M. Lei