基金supported by the National Natural Science Foundation of China(31961143023,31670453)the Strategic Priority Research Program of the Chinese Academy of Sciences(XDA23080302)+2 种基金the National Key Research and Development Program of China(2021YFF0703905)the Chinese Ecosystem Research Network(CERN)the Dinghushan Forest Ecosystem Positioning Research Station of the National Science and Technology Infrastructure Platform and the Operation Service Project of the National Scientific Observation and Research Field Station of the Dinghushan Forest Ecosystem of Guangdong,the Ministry of Science and Technology of the People’s Republic of China.