基金国家自然科学基金(No.91114201)国家重点基础研究发展计划(2012CB821906)资助出版+2 种基金the project CGL2009-11838(Government of Spain)support from the SYNTHESYS Project http://www.synthesys.info/which is financed by European Community Research Infrastructure Action under the FP7 Integrating Activities ProgrammeThe support of the Palaeontology Investment Fund of The Natural History Museum