Vegetation-environment relationships in the forests of Chitral district Hindukush range of Pakistan

We investigated the composition of plant communities to quantify their relationships with environmental parameters in the Chitral Hindukush range of Pakistan. We sampled tree vegetation using the Point Centered Quarter （PCQ） method while understory vegetation was sampled in 1.5-m circular quadrats. Cedrus deodara is the national symbol of Pakistan and was dominant in the sampled communities. Because environmental variables determine vegetation types, we analyzed and evaluated edaphic and topographic factors. DCA-Ordination showed the major gradient as an amalgam of elevation （p〈0.05） and slope （p〈0.01） as the topographic factors correlated with species distribution. Soil variables were the factors of environmental significance along DCA axes. However, among these factors, Mg2＋ , K ＋ and N2＋ contributed not more than 0.054% 0.20% and 0.073%, respectively, to variation along the first ordination axis. We conclude that the principal reason for weak or no correlation with many edaphic variables was the anthropogenic disturbance of vegetation. The understory vegetation was composed of perennial herbs in most communities and was most dense under the tree canopy. The understory vegetation strongly regulates tree seedling growth and regeneration patterns. We recommend further study of the understory vegetation using permanent plots to aid development of forest regeneration strategies.

Vegetation-environment relationship in conifer dominating forests of the mountainous range of Indus Kohistan in northern Pakistan

Environmental variables play a crucial role in shaping vegetation structure,mainly in mountainous ecosystems.Different studies have attempted to identify the environment-vegetation relationship of Conifer Dominating Forests(CDF)worldwide.However,due to differences in local climate and soil composition,different environmental drivers can be found.By applying multivariate analysis techniques,this study investigated the vegetation-environment relationship of CDF of Indus Kohistan in northern Pakistan.Our results showed that CDF of Indus Kohistan are distributed in five distinct ecological groups,which are dominated by different trees and understory species.A total of 7 trees and 71 understory species were recorded from the sampling sites.Cedrus deodara was the leading species among four groups,having the highest importance value(IV),density and basal area.Group I was dominated by Pinus wallichiana with the second highest importance value,density and basal area.In addition,elevation,slope,maximum water holding capacity(MWHC),soil moisture(SM),total organic matter(TOM),sodium,phosphorus and nickel showed highly significant influence on composition and distribution pattern of Indus Kohistan vegetation.Therefore,this study shows a new evidence of vegetation-environment relationship,pointing out specific drivers of vegetation structure in CDF of Indus Kohistan region in northern Pakistan.

Population structure,age and growth rates of conifer species and their relation to environmental variables at Malam Jabba,Swat District,Pakistan

This study focuses on age,growth rate and diameter distribution of pine forests in the Malam Jabba area,Swat District,Pakistan.Wood core samples were taken from twenty stands.Picea smithiana was the oldest at 234 years with a 112-cm diameter.Abies pindrow was 125 years with an 80-cm diameter while the oldest Pinus wallichiana was 122 years with 75-cm diameter.The fastest overall growth rate of 1.5±0.1 year/cm was for P.wallichiana on a west-facing aspect,while the slowest 5.8±2.6 year/cm growth was P.smithiana on an eastfacing exposure.P.wallichiana and A.pindrow exhibited marked differences in growth rates over a 5-year period.The highest growth was by P.wallichiana from 1966 to 2006.A.pindrow showed less growth over the same years,such pattern simultaneously reverse from 1911 to 1965.The relationship between diameter and age,diameter and growth rate and age and growth rate were correlated.P.wallichiana and A.pindrow ages were correlated with diameter and growth rates.P.smithiana age was positive correlated with diameter.Generally,topographic and edaphic factors did not show significant correlations with growth rates,although some appreciable correlations were recorded.The growth of P.wallichiana was correlated with elevation while A.pindrow was correlated with maximum water retaining capacity.Diameter and age produced uneven size classes and many size gaps,which could be the result of anthropogenic disturbances.