Structure and heterogeneity of habitat determine diet of predators despite prey abundance:Similar response in Long-eared,Short-eared Owls and Common Kestrels

According to one of the theses of optimal foraging theory,main prey species abundance in the hunting area is the main factor determining the diet and habitat choices of birds of prey.However other factors can also be important.The habitat structure influences the predators’diets as well.In this study we examined the influence of habitat structure on diet compositions of three species of birds:Long-eared Owl(Asio otus),Short-eared Owl(A.flammeus)and Common Kestrel(Falco tinnunculus).The study was carried out from 2007 to 2019 in a 48 km2area of the Crane’s Homeland Reserve,Moscow Region,Russia.The habitat structures of model species’hunting territories(ratio of different types of landscape elements)were classified in module"Semi-Automatic Classification Plugin"based on the QGIS.A boosted regression tree analysis identified that the share of the main prey species in the diet is primarily determined by the landscape structure of hunting territories.The largest share of Common Vole(Microtus arvalis)in birds’diet was determined by the shrubs area(15%of hunting area),the meadow area(75%),the habitat heterogeneity(70%)and the arable land area(5%).The same predictors determined the largest share of Root Vole(Microtus oeconomus):the shrubs area 25%,the meadow area 70%,and the arable land area 3%.The annual mean abundance of prey species did not determine their importance in the diet of birds of prey.Thus,the main prey abundance in the hunting area is not a determining factor for the formation of diet composition of birds of prey.

Analytical solution of axisymmetric contact problem about indentation of a circular indenter into a soft functionally graded elastic layer

The paper addresses a contact problem of the theory of elasticity,i.e.,the penetration of a circular indenter with a flat base into a soft functionally graded elastic layer.The elastic properties of a functionally graded layer arbitrarily vary with depth,and the foundation is assumed to be elastic,yet much harder than a layer.Approximated analytical solution is constructed,and it is shown that the solutions are asymptotically exact both for large and small values of characteristic dimensionless geometrical parameter of the problem.Numerical examples are analyzed for the cases of monotonic and nonmonotonic variations of elastic properties.Numerical results for the case of homogeneous layer are compared with the results for nondeformable foundation.