基金funded by the Central Government Guides Local Projects of China (XZ202101YD0016C)the open funding from Tibet Joint Key Laboratory of Ecological Security (STAQ-2021T-2)+1 种基金the open funding from Yunnan Key Laboratory of Plant Reproductive Adaption and Evolutionary Ecology,Yunnan University (2019DG056)the National Natural Science Foundation of China (31822007 and 32071660).