基金Financial support provided by the National Key Research and Development Project (2017YFD0101002), the Natural Science Foundation of Shanxi Province (201601 D021128), the Postdoctoral Science Foundation of Shanxi Academy of Agricultural Sciences (YBSJJ1808), the CAAS Innovation Team (CAAS-GJHZ201700X), and the National Engineering Laboratory of Crop Molecular Breeding is gratefully appreciated.