基金Acknowledgments This work was supported by the National Natural Science Foundation of China (31372183, 31000950). We thank the Chinese National Climate Data Center for his- torical climatic data, and the Tibetan government for permis- sion to collect specimens. We thank R. Brown, S. Mullin for their kind detailed reviews on the manuscript before submis- sion, and appreciate R. Brown for his extensive help in im- proving the language of the manuscript before publication. We also thank S. Meiri and another anonymous referee for their kind revisions or suggestions on our submitted manuscript.