基金This research was supported by grants from the National Natural Science Foundation of China(41731175,31770524 and 31872994)the Strategic Priority Research Program A of the Chinese Academy of Sciences(XDA20050101)+1 种基金the Joint Key Research Fund(U20A2005)under cooperative agreement between the National Natural Science Foundation of China(NSFC)and Tibet Autonomous Region(TAR)he Second Tibetan Plateau Scientific Expedition and Research(STEP)program(2019QZKK0302 and 2019QZKK0608).