Vegetation Evolution in the Northern South China Sea Region Since 40 ka BP - An Attempt to Reconstruct Palaeovegetation Based on Biomization

Vegetation evolution in the northern South China Sea region since 40 500 a BP is reconstructed using biomization procedure based on pollen data from deep sea core 17940. The result shows that, it is feasible to reconstruct palaeovegetation using biomization procedure, when pollen, particularly Pinus pollen, transported by mind over long distance is excluded. Results from factor analysis suggest that humidity and temperature are the two main factors determining vegetation evolution on land around the northern South China Sea. From 40 500 a BP to 11 100 a BP, broad-leaved evergreen forest (WAMF), and montane conifers(MGRF) occurred on hills and low mountains; while steppe (STEP) predominated on the exposed shelf. The main feature of the vegetation evolution is the frequent alternation between MGRF (or WAMF) and STEP, implying abrupt changes in millennium scale between humid/cold and dry/temperate climate. All abrupt climate events could be broadly correlated with Henrich events and Dansggard-Oscherge events. One of the events around 12 700 a BP, sees the occurrence of MGRF, suggesting that climate turned humid and cold rapidly. This may be correlated with the Younger Dryas event; Broad-leaved evergreen (WAMF) predominates since 11 000 a BP. During the early Holocene and late Holocene tropical rainforest (TRFO) or tropical seasonal forest (TSFO) occurred several times.

Palaeovegetational and palaeoenvironmental changes since the last deglacial in Gonghe Basin, northeast Tibetan Plateau

Paleoenvironmental history in the monsoonal margin in the northeast Tibetan Pla- teau provides important clue to the regional climate. Previous researches have been limited by either poor chronology or low resolution. Here we present a high-resolution pollen record from a 40.92-m-long sediment core （DLH） taken from Dalianhai, a terminal lake situated in the Gonghe Basin, the northeast Tibetan Plateau for reconstructing the vegetation and climate history since the last deglacial on the basis of a chronology controlled by 10 AMS 14C dates on plant remains preserved in the core sediments. The pollen assemblages in DLH core can be partitioned into 6 pollen zones and each zone is mainly characterized by the growth and decline of tree or herb pollen percentage. During the periods of 14.8-12.9 ka and 9.4-3.9 ka, the subalpine arboreal and local herbaceous pollen increased, indicating the subalpine forest developed in the surrounding mountains and a desert steppe or typical steppe developed in Gonghe Basin under a relatively moister climate. During the periods of 15.8-14.8 ka, 12.9-9.4 ka and 3.9-1.4 ka, the forest shrank or disappeared according to different degrees of aridity, and the desert steppe degraded to a more arid steppe desert in the basin, indicating a dry climate. After 1.4 ka, vegetation type around Dalianhai was mainly dominated by steppe suggested by increased Artemisia. Our results suggested the climate history in this region was dry from 15.8-14.8 ka, humid from 14.8-12.9 ka and dry from 12.9-9.4 ka, after which the climate was humid during 9.4-3.9 ka, followed by dry conditions during 3.9-1.4 ka and humid conditions in the last 1.4 ka. The change of pollen percentage and the evolution of palaeovegetation in Dalianhai since the last deglacial were similar to those recorded in Qinghai Lake. The forest expanded in the mountains around Dalianhai during the Bol- ling-Aller^d period, shrank during the Younger Dryas and the early Holocene, then it devel- oped and reached its maximum in the mid-Holocene. During the late Holocene, the vegetation began to shrink till disappearance. However, the timing of forest expansion in the Holocene lagged behind that of Qinghai Lake, and this spatial heterogeneity was probably caused by the different forest species between these two places. The maximum of forest development in the mid-Holocene was inconsistent with the period of stronger summer monsoon in the early Holocene indicated by stalagmite records, the reason might be related to the complexity of vegetation response to a large-scale climatic change.

Temporal-Spatial Distribution of Vegetation in the Qinghai-Xizang Plateau During the Past 12 ka BP

Comprehensive studies on the basis of pollen records from lake cores at 30 sites in the Qinghai-Xizang Plateau have been used to reconstruct temporal-spatial distributions of Holocene vegetations. Before the, Holocene (prior to 12.0 ka BP) desert steppe vegetation was developed from the, east to the west in the most parts of the Plateau, with a few exceptions in the extreme southeastern appeared. During the early Holocene ( 12.0 - 9.0 ka BP) deciduous broad-leaved forest/conifer and broad-leaved deciduous mixed forest were distributed in the east of Plateau (104degrees - 98degrees E). Meadows or shrub, meadow appeared in the middle of tire Plateau (98degrees - 92degrees E). Farther west to about 80degrees E, a steppe landscape was present. During the middle Holocene (9.0 - 3.2 ka BP), the palaeovegetations were sequentially conifer and broad-leaved deciduous mixed forest and sclerophyllous broad-leaved forest (104degrees - 98degrees E) - conifer and broad-leaved deciduous mixed forest (98degrees - 94degrees E) - shrub meadow (94degrees - 92degrees E) - steppe (92degrees - 80degrees E). During the late Holocene (after 3.2 ka BP), the palaeovegetations were sequentially sclerophyllous broad-leaved forest - conifer and broad-leaved deciduous mixed forest - meadow - steppe - desert from east to west of the Plateau.

Siwalik plant megafossil diversity in the Eastern Himalayas: A review

The Eastern Himalayas are renowned for their high plant diversity.To understand how this modern botanical richness formed,it is critical to investigate past plant biodiversity preserved as fossils throughout the eastern Himalayan Siwalik succession(middle Miocene-early Pleistocene).Here,we present a summary of plant diversity records that document Neogene floristic and climate changes.We do this by compiling published records of megafossil plant remains,because these offer better spatial and temporal resolution than do palynological records.Analyses of the Siwalik floral assemblages based on the distribution of the nearest living relative taxa suggest that a tropical wet evergreen forest was growing in a warm humid monsoonal climate at the deposition time.This qualitative interpretation is also corroborated by published CLAMP(Climate Leaf Analysis Multivariate Program) analyses.Here,we also reconstruct the climate by applying a new common proxy WorldClim2 calibration.This allows the detection of subtle climate differences between floral assemblages free of artefacts introduced by using different methodologies and climate calibrations.An analysis of the Siwalik floras indicates that there was a gradual change in floral composition.The lower Siwalik assemblages provide evidence of a predominance of evergreen elements.An increase in deciduous elements in the floral composition is noticed towards the close of the middle Siwalik and the beginning of the upper Siwalik formation.This change reflects a climatic difference between Miocene and Plio-Pleistocene times.This review helps us to understand under what paleoenvironmental conditions plant diversity occurred and evolved in the eastern Himalayas throughout the Cenozoic.

Aliphatic biomarker signatures of early Oligocene-early Miocene source rocks in the central Qiongdongnan Basin:Source analyses of organic matter

The geochemical signatures of fifty-four rock samples and three supplementary drill stem test(DST)oils from the Yacheng-Sanya formations in the central Qiongdongnan Basin(CQB)were analysed.Reconstruction of the early Oligocene-early Miocene(36–16 Ma)palaeovegetation and source analyses of organic matter(OM)were conducted using aliphatic biomarkers in ancient sediments and DST oils.Both the interpreted aquatic and terrigenous OM contributed to the CQB source rocks(SRs)but had varying relative proportions.The four distribution patterns derived from n-alkanes,terpanes,and steranes are representative of four OM composition models of the Yacheng-Sanya SRs,including model A,model B,model C,and model D,which were classified based on the increasing contribution from terrigenous OM relative to aquatic OM.Some terrigenous higher plantderived biomarkers,including oleanane,des-A-oleanane,C_(29)ααα20R sterane,bicadinanes,the C_(19)/(C_(19)+C_(23))tricyclic terpane ratio,and other n-alkane-derived ratios suggest that angiosperms had increased proportions in the palaeoflora from early Oligocene to early Miocene,and the bloom of terrigenous higher plants was observed during deposition of upper Lingshui Formation to lower Sanya Formation.These findings are consistent with the incremental total organic carbon and free hydrocarbons+potential hydrocarbons(S_1+S_2)in the lower Lingshuilower Sanya strata with a significant enrichment of OM in the E_3l_1-N_1s_2 shales.The maturity-and environmentsensitive aliphatic parameters of the CQB SRs and DST oils suggest that all the samples have predominantly reached their early oil-generation windows but have not exceeded the peak oil windows,except for some immature Sanya Formation shales.In addition,most of the OM in the analysed samples was characterised by mixed OM contributions under anoxic to sub-anoxic conditions.Furthermore,terrestrial-dominant SRs were interpreted to have developed mainly in the Lingshui-Sanya formations and were deposited in sub-oxic to oxic environments,compared to the anoxic to sub-anoxic conditions of the Yacheng Formation.

Climate characteristics of the eastern Mongolian Plateau,China during the early Early Cretaceous(145-132 Ma):Palynological evidence from the Tongbomiao Formation in Well Hong-6,Hailar Basin

This study identified two palynological assemblages,namely Bayanhuasporites-Cycadopites-Protoconiferus and Cicatricosisporites-Cedripites-Perinopollenites,in the Tongbomiao Formation in the Hongqi Sag in the Hailar Basin,Inner Mongolia,China for the first time.The former is distributed in the lower part of the Tongbomiao Formation and is characterized by abundant gymnosperm pollen and diverse fern spores.Among them,the gymnosperm pollen is dominated by Paleoconifer(4.98%-31.62%)and Cycadopite(8.55%-25.23%)pollen grains and also includes other pollen grains such as Classopollis,Parcisporites,Erlianpollis,Callialasporites,and Jiaohepollis.The fern spores in the former palynological assemblage contain Bayanhuasporite(0-8.96%),Granulatisporites(0.93%-6.97%),and some important Cretaceous genera,such as Cicatricosisporites,Concavissimisporites,Densoisporites,Hsuisporites,Foraminisporis,and Leptolepidites.The Cicatricosisporites-Cedripites-Perinopollenites palynological assemblage is distributed in the upper part of the Tongbomiao Formation.Gymnosperm(77.30%),Pinaceae(31.9%),and Paleoconiferus(19.02%)pollen predominate this palynological assemblage,and Quadraeculina,Erlianpollis,and Jiaohepollis pollen are also common in this assemblage.The fern spores in this palynological assemblage include abundant Cicatricosisporites(4.29%).Besides,Concavissimisporites,Aequitriradites,and Leptolepidites are also common in this palynological assemblage.No angiosperm pollen has been found in both palynological assemblages.The identification of both palynological assemblages provides important evidence for the biostratigraphic correlation between the Hailar Basin and its adjacent areas.It also enables the reconstructions of the Berriasian-Valanginian(Early Cretaceous)vegetation and the paleoclimate on the eastern Mongolian Plateau during 141-132 Ma.The vegetation reconstructed on the palynological data of the represented by Hailar Basin in eastern Mongolian Plateau(141.6-141.4 Ma),form conifer forest or conifer broad-leaved mixed forest to conifer forest with shrubs and grassland,the climate belongs to warm temperate and warm-subtropicalt,the highest temperature is estimated to reach 35-38℃.Form 132.3 Ma,the vegetation type is conifer forest,and its paleoclimate is sub-humid warm temperate,the highest temperature is estimated to reach 24-29℃.

Palynological evidence for vegetational and climatic changes from the HQ deep drilling core in Yunnan Province, China

The high-resolution pollen study of a 737.72-m-long lake sediment core in the Heqing Basin of Yunnan Province shows that the vegetation and climate of mountains around the Heqing Basin went through six obvious changes since 2.780 Ma B.P. Namely, Pinus forest occupied most mountains around the studied area and the structure of vertical vegetational belt was simple between 2.780 and 2.729 Ma B.P., reflecting a relatively warm and dry climate. During 2.729―2.608 Ma B.P., the areas of cold-temperate conifer forest (CTCF) and Tsuga forest increased and the structure of vertical vegetational belt became clear. According to percentages of tropical and subtropical elements growing in low-altitude regions rifely increased, we speculate that the increase of CTCF and Tsuga forest areas mainly resulted from strong uplift of mountains which provided upward expanding space and growing condition for these plants. Thus, the climate of the low-altitude regions around the basin was relatively warm and humid. Between 2.608 and 1.553 Ma B.P., Pinus forest occupied most mountains around the studied area and forest line of CTCF rose, which reflects a moderately warm-dry climate on the whole. During 1.553―0.876 Ma B.P., the structure of vertical vegetational belt in mountains around the studied area became complicated and the amplitude of vegetational belts shifting up and down enlarged, which implies that the amplitude of climatic change increased, the climatic associational feature was more complex and the climate was moderately cold at a majority of the stage. During 0.876―0.252 Ma B.P., there were all vertical vegetational belts existing at present in mountains around the studied area. The elements of each belt were more abundant and complex than earlier. At different periods in the stage vertical vegetational belts occurred as expanding or shrinking, and alternated each other. The amplitude of vegetational belts shifting up and down was the maximum in the whole section. This change suggests that the amplitude of climatic change was evidently larger than earlier, but the frequency reduced and the climatic associational feature was more complex. From 0.252 Ma B.P. to the present, the most time was characteristic of the expanding of Pinus forest and semi-humid evergreen broad-leaved forest (SEBF) in mountains around the studied area, while expanding time of other vegetational belts was very short, which reflects a smaller amplitude of cold and warm fluctuation. On the basis of the six obvious cycles of vegetational and climatic changes, there were still many times of secondary vegetational successions and climatic oscillations. Based on the above analysis, the forcing mechanism of vegeta-tional succession and climatic change in the Heqing Basin is further discussed. It is primarily consid-ered that main influential factors were exterior factors such as orbital parameters, etc., but the uplift of the Qinghai-Tibet Plateau played a very important function for environmental changes in the Heqing Basin at two times obvious increase of vertical vegetational belts and three climatic transitions.

Pedogenic carbonate isotope record of vegetational evolution since late Miocene in Loess Plateau

lsotopic analyses for paleovegetational evolution have been carried out on samples of the pedogenic carbonate nodules in the Red Clay-Loess sequence at Lingtai （35°N）, the Loess Plateau. Stable carbon isotopic composition indicates that （ ⅰ ） C4 plants might be present at least by7.0 Ma; （ ⅱ ） C4 plants expanded gradually between ～4.0 and ～3.2 Ma, and their biomass fraction was up to 50%; and （ ⅲ ） the biomass of C4 vegetation since ～2.0 Ma seems to have been decreased to the level （about 20%） before 4.0 Ma. C4 plant expansion at Lingtai cannot be fully understood with the 'global C4 expansion' model because it occurred much later up to 3.0 Ma than in Pakistan, which indicates that some changes in the regional climatic system may also contribute to C3/C4 shift except changes in atmospheric CO2 concentrations and temperature. The latitudinal zone for C3/C4 transition seems to move southwards slightly in East Asia, compared with the case in North America where 37°N is the ideal

Late Miocene wood flora associated with the Yuanmou hominoid fauna from Yunnan, southwestern China and its palaeoenvironmental implication

The Upper Miocene Xiaohe Formation of the Yuanmou Basin in Yunnan Pro- vince, southwestern China, is famous for its hominoid fauna and is important for studying the Late Cenozoic human and mammal evolution. Abundant fossil wood was found associated with this fauna, which provided important evidence for palaeoenvironmental reconstruction of the basin. Among the fossil wood, two different taxa have been identified namely, Quercoxylon sp. （Fagaceae） and Pterocarya sp. （Juglandaceae）. Based on the habitats of their Nearest Living Relatives （NRLs）, it is suggested that upland subtropical evergreen broad-leaved forest dominated by QuercuslLithocarpus was common around the basin, while a lowland deciduous broad-leaved forest dominated by Pterocarya was present on the river bank of the basin during the Late Miocene.