Variations of plant life form diversity along the Northeast China Transect and its direct gradient analysis

Plant life form diversity and its direct gradient analysis on a larger scale climate change gradient were tested, based on the data from Northeast China Transect platform. The results showed that the species numbers, life form richness and life form diversity were relative higher at the eastern forests and the ecotone between typical vegetation, while those on the meadow grasslands and typical steppes were lower. Although plant life forms can reflect the climate variations, life form diversity is not consistent with the major global gradient along the NECT.

Life form and geographical distribution of plants in Posthband region,Khonj,Fars Province,Iran

A study was conducted to identify plant species,diversity,life form and relevant species to Mediterranean climate,Irano-Torunian and Sahara-Sindian in Postband region（N 27°58＇4＂-28°2＇16＂,E 53°17＇34＂-53°22＇30＂）.In total,95 species belonging to 29 families and 69 genera of flora were surveyed and identified from December 2006 to June 2008.Composite and Gramineae were important families in terms of species frequency.According to Raunkiaer＇s system and using X^2 tests,the life-form spectrum showed that therophytes accounted for 47% of all species,and hemicryptophytes for 31%,phanerophytes for 12%,chamaeophyte for 7%,and cryptophytes for 3%.In geographical distribution,29% species with the most frequency belonged to Irano-Torunian region.Results show that therophytes were more than normal spectrum and phaneropytes were less than normal spectrum,which was in agreement with data obtained in arid climate.

Ecological biomass allocation strategies in plant species with different life forms in a cold desert, China

Biomass allocation patterns among plant species are related to their adaptive ecological strategies. Ephemeral, ephemeroid and annual plant life forms represent three typical growth strategies of plants that grow in autumn and early spring in the cold deserts of China. These plants play an important role in reducing wind velocity in the desert areas. However, despite numerous studies, the strategies of biomass allocation among plant species with these three life forms remain contentious. In this study, we conducted a preliminary quadrat study during 2014–2016 in the southern part of the Gurbantunggut Desert, China, to investigate the allocation patterns of above-ground biomass(AGB) and below-ground biomass(BGB) at the individual level in 17 ephemeral, 3 ephemeroid and 4 annual plant species. Since ephemeral plants can germinate in autumn, we also compared biomass allocation patterns between plants that germinated in autumn 2015 and spring 2016 for 4 common ephemeral species. The healthy mature individual plants of each species were sampled and the AGB, BGB, total biomass(TB), leaf mass ratio(LMR) and root/shoot ratio(R/S) were calculated for 201 sample quadrats in the study area. We also studied the relationships between AGB and BGB of plants with the three different life forms(ephemeral, ephemeroid and annual). The mean AGB values of ephemeral, ephemeroid and annual plants were 0.806, 3.759 and 1.546 g/plant, respectively, and the mean BGB values were 0.106, 4.996 and 0.166 g/plant, respectively. The mean R/S value was significantly higher in ephemeroid plants(1.675) than in ephemeral(0.154) and annual(0.147) plants. The mean LMR was the highest in annual plants, followed by ephemeroid plants and ephemeral plants, reflecting the fact that annual plants allocate more biomass to leaves, associated with their longer life span. Biomass of ephemeral plants that germinated in autumn was significantly higher than those of corresponding plants that germinated in spring in terms of AGB, BGB and TB. However, the R/S value was similar in plants that germinated in autumn and spring. The slope of regression relationship between AGB and BGB differed significantly among the three plant life forms. These results support different biomass allocation hypotheses. Specifically, at the individual level, the AGB and BGB partitioning supports the allometric hypothesis for ephemeroid and annual plants and the isometric hypothesis for ephemeral plants.

Antioxidant systems of aquatic macrophytes in three life forms:a case study in Lake Erhai,China

Antioxidant systems are vital in life activities of macrophytes. Species with diff erent life forms need to cope with distinct environments by modifying physiological characters, especially antioxidant systems. In order to find diff erences among life forms and consequence of lake eutrophication, we studied three antioxidant enzymes activity (superoxide dismutase (SOD), ascorbate oxidase (APX) and catalase (CAT)) and total soluble phenolics (TP) content in leaves of 26 macrophyte species in September 2013 in Lake Erhai, China. We found that antioxidation varied accordingly with life forms. The activities of SOD and APX in emergent macrophytes (EM) and floating-leaved macrophytes (FM) were much lower than those of submerged macrophytes (SM). On the contrary, TP content was much higher in EM and FM species. There was a negative correlation between TP and antioxidant enzyme activities (CAT and APX). The results suggested that EM and FM species rely on phenolics might to adapt to adverse environments (higher herbivores predation pressure and UV radiation intensity), while SM species more rely on antioxidant enzymes possibly due to lower demand for antioxidation and/or lack of light and inorganic C availability for phenolics synthesis. We also found FM species represent highest fitness in term of antioxidant system, which would lead to overgrowth of FM species and littoral zone bogginess during lake eutrophication. Finally, it is necessary to carry out the verification experiment under the control condition in the later stage, especially for the dominant ones in eutrophic lakes, to understand the exact adaptive mechanisms of them.

Characteristics of life-form and growth-form of plant species in an alpine ecosystem of North-West Himalaya

The present study was conducted in the alpine pastures of Tungnath （30° 14＇ N and 79° 13＇ E） to observe life-form and growth-form patterns of alpine plant species under grazed and ungrazed conditions and to work out the plant life form spectrum. Species were categorized as plant habit, height and length of growth-cycle and life-form classes according to Raunkiaer＇s system. The results show that in total of 68 species at grazed site, hemicryptophytes （He） accounted for 50.00% species, followed by cryptophytes （26.47%）, chamaephytes （16.18%）, phanemphytes （4.41%） and therophytes （2.94%）. At the ungrazed site in 65 plant species, hemicryptophytes （He） accounted fo＇r 49.23% species, cryptophytes （26.15%）, chamaephytes （15.38%）, phanerophytes （6.15%） and ther6phytes （3.08 % species）. In general, hemicrptophyte are dominant^in both sites i.e. graged and ungrazed. Growth form categories were classified as forbs, shrubs, grasses and sedges＇and undershrubs, according to plant habit and height. On the basis of length of the growth cycle, species were categorized as plant species of short growth cycle, intermediate growth cycle and long growth cycle. The short forbs of plant habit and height, had the highest emergence, and grasses and sedges had the lowest emergence in representative species. Percentage of species with long growth cycle was highest in both sites.

Functional Enrichment of Utopian Distribution of Plant Life-Forms

Adaptive fuzzy neural inference systems are used to illustrate the primary nodal number of plant life-forms. Categorization of two candidate areas is carried out using the water-energy dynamic (for Ecuador, South America) and Macedonia, Southern Europe), within which the life-form spectra are distributed. Genetic optimization methods are used to expand the primary nodal number to the complete number of life-form categories. The distribution of the elements exhibits a stochastic, binomial distribution and the utopia line and curve are summarized which enhance accuracy of the climatic data and of the consequent numbers of plant species occurrences. Expansion of the distribution of each life-form category is approximated within the Z utopia hyperplane with use of the functional approximation algorithm. This process gives additional structure and informative value to the Z plane, enhancing our ability to make informed policy decisions concerning species and ecosystem conservation.

2011-2015年洞庭湖洲滩群落植物生态型、生活型和生长型组成数据集

植物群落结构与环境的关系可以通过研究植物生活型和生态型进行深入的解析,同时植物生活型是研究植物群落的发生、发展及演替规律的重要内容。中国科学院洞庭湖湿地生态系统观测研究站按中国生态系统研究网络(CERN)监测指标体系,在洞庭湖周期性水文变化情势下,对洞庭湖湿地洲滩典型植物群落类型的物种组成进行长期定位监测。通过专家对洞庭湖湿地3个典型植物群落(苔草、南荻和水蓼)的长期监测数据集进行验证,确保数据的准确性,然后进行分类处理,得到洞庭湖洲滩2011-2015年植物群落植物生活型、生长型和生态型数据集。本数据集包含有植物种名、拉丁名、生活型、生长型、光生态型、水分生态型、土壤pH值生态型、淤积生态型,共8个指标,同时提供了完整的样地信息和样地管理记录,为数据的科学利用提供了空间和时间上的背景信息。本数据集为分析洞庭湖湿地洲滩植被群落生态型和生活型组成,植物群落与环境的相关提供了数据支撑,是深入研究洞庭湖湿地植物被演替的重要参考依据。

金马河沿岸次生林苔藓植物群落对人为干扰的响应

为探索人为干扰对四川金马河沿岸次生林苔藓植物群落的影响,通过分析比较不同干扰程度(轻度、中度、重度)区域苔藓植物的生态位、多样性、生活型、生长附着基质,研究苔藓植物群落特征和物种分布规律。结果表明,在金马河沿岸次生林地12块样地中共调查到苔藓植物15科23属34种,其中,苔类植物3科4属4种,藓类植物12科19属30种。优势科主要为青藓科和羽藓科,其中,细叶小羽藓具有最大的重要值和生态优势度,其重要值、生态位宽度和生态优势度分别为0.5368、2.4648、0.1169,占据重要生态位,台湾青藓次之。耐旱性强的丛集型苔藓植物种类随着人为干扰程度的增强而增加,占比从30.43%逐渐增至40.91%,且其重要值在重度干扰区位于前列。重度干扰区苔藓植物群落的物种丰富度指数和Shannon-Wiener多样性指数均显著低于轻、中度干扰区(P<0.05),中度干扰区苔藓植物群落的Pielou均匀度指数显著高于其他两个干扰区(P<0.05)。重度干扰区苔藓植物附着在腐(枯)木的数量显著少于轻度干扰区(P<0.05),附着在树干的数量显著少于中度干扰区(P<0.05),且主要附着在土壤和岩石上,附着基质逐渐倾向于单一类型。金马河沿岸次生林重度干扰区的苔藓植物物种丰富度指数、Shannon-Wiener多样性指数显著降低(P<0.05),过度干扰会导致林地内生态环境和苔藓植物群落结构遭受破坏。

人类文明形态演进的生命辩证法

马克思的世界观实现了相对于旧哲学世界观的革命性变革,这就是把价值论的维度引入世界观,从而不仅回答了“世界是什么”的本体论问题,而且回答了“世界向何处去”的价值论问题。按照马克思的世界观,人类文明形态的演进本质上是人类这一物种寻求永生的过程。于是,世界历史进程便表现为人类文明形态不断演进的过程。文明形态的演进遵循辩证法原理,而人类通过文明形态的演进,展现了人类独特的生命活动形式,即生命辩证法。共产主义社会文明形态实现了人类文明中“自然法则”和“理性法则”的辩证统一,使人类这一物种的永久生存成为可能。

Host plant traits play a crucial role in shaping the composition of epiphytic microbiota in the arid desert,Northwest China

Phyllosphere microorganisms are a crucial component of environmental microorganisms,highly influenced by host characteristics,and play a significant role in plant health and productivity.Nonetheless,the impact of host characteristics on shaping phyllosphere microbial communities of plants with different life forms remains ambiguous.Utilizing high-throughput sequencing technology,this study analyzed the diversity and community composition of phyllosphere epiphytic microorganisms(e.g.,bacteria and fungi)of various plant life forms in the hinterland of the Gurbantunggut Desert,Northwest China.Functional annotation of prokaryotic taxa(FAPROTAX)and fungi function guild(FUNGuild)were employed to assess the ecological functions of microorganisms and to investigate the role of stochastic and deterministic processes in shaping phyllosphere microbial communities.Result showed a diverse array of phyllosphere epiphytic microorganisms in the desert plants,with Proteobacteria,Cyanobacteria,and Actinobacteriota dominating bacterial community,while Ascomycota and Basidiomycota were prevalent in fungal community.Comparison across different plant life forms highlighted distinct microbial communities,indicating strong filtering effects by plant characteristics.FAPROTAX prediction identified intracellular parasites(accounting for 27.44%of bacterial community abundance),chemoheterotrophy(10.12%),and phototrophy(17.41%)as the main functions of epiphytic bacteria on leaves of different life form plants.FUNGuild prediction indicated that phyllosphere epiphytic fungi primarily served as Saprotrophs(81.77%),Pathotrophs(17.41%),and Symbiotrophs(0.82%).Co-occurrence network analysis demonstrated a predominance of positive correlations among different microbial taxa.Raup-Crick dissimilarity index analysis revealed that deterministic processes predominantly influenced phyllosphere bacterial and fungal community assembly.Variance partitioning analysis and random forest modeling suggested that plant leaf functional traits significantly impacted both bacterial and fungal community composition,with fungal community composition showing a closer association with leaf nutrients and physiology compared with bacterial community composition.The distinct responses of bacterial and fungal communities to plant traits were attributed to the differing properties of bacteria and fungi,such as bacteria having higher potential dispersal rates and broader ecological niches than fungi.Overall,the results indicate that phyllosphere bacterial and fungal communities undergo similar community assembly processes,with fungi being more influenced by plant characteristics than bacteria.These findings offer novel insights into the ecology of phyllosphere microbial communities of desert plants.

中国省级重点保护野生植物的种类组成和多样性分析

为揭示我国省级重点保护野生植物的主要特征,对我国已公布省级重点保护野生植物名录的24个省份数据进行了整理,并初步分析其种类组成、生活型谱和濒危等级。结果显示:(1)我国省级重点保护野生植物共有2044种,隶属于208科,844属。其中,大型真菌有23种,苔藓植物有14种,石松类和蕨类植物有63种,裸子植物有50种,被子植物有1894种。不同省份的重点保护植物种数介于17~448种,平均种数为115种。(2)植物的生活型共有10种。其中以草本植物最多,有1200种,占总种数的58.71%;其次为常绿乔木,有262种,占总种数的12.82%;再次为落叶乔木,有225种,占总种数的11.01%。(3)根据世界自然保护联盟(IUCN)红色名录的濒危等级划分,这2044种植物可分为6种类型。其中以无危(LC)最多,有1224种,占比59.88%;其次为易危(VU),有248种,占比12.13%;再次为近危(NT),有198种,占比9.69%;濒危(EN)有173种,占比8.46%;极危(CR)有128种,占比6.26%。值得注意的是,濒危植物类群(包括CR、EN、VU、NT)的植物合计有747种,占比36.55%。研究发现,我国省级尺度的重点保护野生植物名录涵盖的植物类群广泛,但是各省制定的名录涉及的植物种数和比例差异很大。不同省份的重点保护野生植物的生活型多样,但是总体上以草本植物为主;这些植物的濒危等级相对较高,这可能与其地理分布相对狭窄、种群规模往往较小以及受到一定人为活动的影响有关。因此,建议国家及省级相关职能部门在省级尺度上对重点保护野生植物名录制定加强指导,在强调物种的珍稀、濒危和特有性的同时,更好地兼顾管理的可操作性、有效性和充分性。

拉埃尔·耶吉异化与生活形式批判理论视域下的家庭

拉埃尔·耶吉关于家庭的思考是她的批判理论中非常重要的一个主题,这既是她的生活形式批判的例证之一,又是她个人重新确定家庭本性的尝试。耶吉重新激活并改写的“异化”理论和“生活形式”的理论是她对资本主义实践情景中的家庭做出批判性思考并引导批判理论的当代话语转向的前提。她通过无力且失控的生活、角色行为、内部分裂和冷漠四种范例考察了异化视域下的家庭生活形式本身。耶吉把家庭看作由社会实践的文化秩序生发出来的具象生活形式,以此为基础建构了一种生活形式本体论,阐释家庭何以能成为合适的批判对象,同时可以应用于解决问题的实例。她从问题规范层面的家庭、黑格尔对资产阶级家庭的伦理解释是解决问题的尝试、特定家庭形式问题的规范性失败和家庭问题的内在批判等四个方面就家庭问题做了深入的剖析,从异化的角度出发构建出一种家庭批判观。耶吉的家庭观既在一定程度上误解了黑格尔的家庭观,又有可能陷入把家庭当作私人事务而忘记了马克思主义的本性的危险。

明堂山主要药用植物资源调查分析

为全面了解岳西县明堂山药用植物资源的组成类型、生活型和药用部位等资源现状,本文采用传统野外调查方法收集药用植物资源的分布信息,通过形态观察及查阅相关文献资料对药用植物进行物种鉴定和药用植物整理分析.结果表明,本次调查发现药用植物231种,隶属96科205属,以小型科(2~4种)和单种科(1种)为主.生活型以草本植物占绝对优势(61.47%);药用部位以全草类、根及根茎类占多数(61.48%);珍稀濒危药用植物29种,其中国家一级保护植物2种,二级保护植物8种.岳西县明堂山药用植物资源类型多样性特征明显,生态环境保护良好,此次调查为后续有效保护和合理开发药用植物资源提供科学依据.

脑机接口、生命形式与同一性

脑机接口技术能够修复与增强肢体功能,却带来了技术中断的问题,即导致了个体同一性的中断。但是对脑机接口的同一性进行反思需要克服现代思维的束缚,这一思维建立在生命连续性的基础上。而从神话思维角度看,生命中断是本真的生命现象。从这一观念看,脑机接口导致的中断担忧可以被舒缓。脑机接口导致的技术中断依然存在,确实对生命整体性构成损害;但是其通过功能训练实现了身体功能的延续,这值得肯定。

西北干旱区荒漠植物叶片和细根碳、氮、磷化学计量特征

植物的叶片和细根是植物地上部分和地下部分最重要的营养器官,其碳(C)、氮(N)、磷(P)化学计量特征反映了植物的养分限制状况及叶片与细根间的协同作用,开展叶片与细根化学计量关系的研究,对认识植物养分利用策略及元素间平衡关系有重要意义。对西北干旱区21种荒漠植物叶片与细根C、N、P含量进行了测定,分析了C、N、P含量及其计量比在不同生活型及不同器官间的关系。结果表明:1)西北干旱区荒漠植物叶片与细根C含量相对稳定,叶N、P含量高于细根,说明在干旱生境中,植物对养分的分配侧重于地上部分,以完成其正常的生命活动;叶片与细根的N∶P无显著差异,有明显的保守性,表明植物地上与地下部分养分吸收策略与分配规律具有一致性;2)不同生活型植物叶片和细根的C、N、P含量及C∶N、C∶P存在显著差异,灌木与禾草具有较高的C∶N和C∶P,杂类草具有较高的N、P含量,说明灌木与禾草倾向于保守型养分适应策略,杂类草倾向于快速生长的资源竞争策略;3)21种荒漠植物叶片与细根N、P含量间均存在显著的正相关关系,表明西北干旱区荒漠植物体内N、P元素间存在相互作用;4)植物叶片与细根间C、N、P含量及C∶N、C∶P存在等速生长关系,表明植物光合产物和养分在地上与地下部分间的分配具有平行的比例关系,但这种关系受生活型影响。上述结果表明西北干旱区不同生活型植物在恶劣生境中存在不同的养分适应策略,且地上与地下部分的养分利用策略及分配规律存在协同作用,这为进一步认识西北干旱区荒漠植物的养分利用策略提供了试验基础。

大尺寸均质化M50轴承钢坯的构筑成形

针对大尺寸M50轴承钢锭中液析碳化物粗大的问题,提出采用金属构筑成形技术制备大尺寸均质化M50轴承钢坯的新思路。通过基元加工、真空封焊、高温变形、保温扩散和多向锻造工艺制备了80 mm×80 mm×1000 mm的M50轴承钢构筑坯料。拉伸和疲劳性能的试验结果表明,M50轴承钢构筑界面实现了完全冶金结合,其力学性能达到与基体相当的水平,通过构筑成形技术制备的均质化M50轴承钢坯的疲劳性能显著优于采用一体化冶炼工艺制备的大尺寸M50轴承钢坯。

塔里木盆地南缘绿洲-沙漠过渡带8种植物C、N、P生态化学计量特征的季节变化

研究荒漠生态系统中乔木、灌木和草本叶片生态化学计量特征及其随季节变化的规律,旨在阐明不同生活型植物生长发育过程中生态化学计量特征动态平衡及其差异,为荒漠生态系统的保护与管理提供理论依据。选择塔里木盆地南缘绿洲-沙漠过渡带内不同生活型8种植物,在生长季的4—9月连续进行叶片采集,分析叶片C、N、P含量及其计量比在整个生长季的变化。结果表明:(1)整体上,8种植物叶片C含量的季节变化不大,N、P含量在生长初期最高,之后逐步下降,体现了明显的稀释作用,而植物叶片的C∶N、C∶P和N∶P在整个生长季内的变化规律不一致;(2)综合分析8种植物叶片C、N、P含量及其计量比值在整个生长季节内的变异,变异系数由大到小排序为:P(32.39%)>C∶P(25.67%)>N(20.96%)>N∶P(20.42%)>C∶N(19.83%)>C(6.49%);(3)植物叶片C含量的变异主要受物种和生活型的影响,N、P含量及C∶N、C∶P的变异受物种和生长季的影响最大,而N∶P的变异受到生长季与物种的交互影响大于生长季的影响;(4)8种植物叶片N、P含量间存在极显著的正相关关系(P<0.001);P含量则与N∶P间呈现显著负相关(P<0.001),而N含量与N∶P间无显著相关性(P>0.05)。绿洲-沙漠过渡带8种植物叶片碳、氮、磷含量及其化学计量比在生长季因物种而不同,它们在生长季内的变异系数在物种间也存在差异,植物生长更容易受到N元素的限制。

云南思茅松林植物区系、生活型与叶相特征分析

云南的思茅松林为我国云南省重要的战略森林资源,但其植物资源多样性未见系统报道,为揭示其区系特征及群落外貌,本研究基于144个群落调查样方数据,分析了云南的思茅松林的维管植物区系、生活型与叶相特征。结果显示:(1)云南的思茅松林群落维管植物经合计共130科440属800种,其中包括蕨类植物18科36属61种,裸子植物2科4属7种,被子植物110科400属732种,优势科为豆科、壳斗科和禾本科,优势属为榕属、锥属和柯属。(2)科属的地理成分均以热带成分占优势,且均以泛热带成分为主;温带分布型以北温带及其变型为主。未见中国特有科,中国特有属9种。寡种和单种科属占比高,一定程度表明了该群落类型对生境选择的多样化响应。从演化趋势来看,古老成分与新生类群共存。(3)生活型谱中高位芽植物为该群落优势生活型,占比为61.25%,Raunkiaer生活型谱呈“L”型。(4)植物叶相以纸质(44.00%)、单叶(74.50%)、全缘(67.75%)、中型叶(46.38%)为主。该研究结果有助于进一步认识思茅松林区系特征、为其生态保护和管理提供参考价值。

张家口崇礼区森林生物多样性及其垂直分布格局

选择张家口崇礼区东部山林,沿海拔高度设置样地,开展森林植被组成结构调查,分析其植被特征和生物多样性沿海拔高度的垂直分布规律.结果表明:森林植被类型以白桦为优势种的落叶阔叶林为主,随着山地海拔升高,逐渐演变为以华北落叶松为优势种的针叶林和针叶阔叶混交林.森林木本植物的物种多样性沿海拔高度呈现出先升高、后降低的变化趋势,多样性最大值区域位于海拔1900 m;森林物种丰富度主要受灌木层物种变化的影响.森林地表枯落物在海拔1500 m和1900 m样地积累较多,乔木植物对地表枯落物的贡献较大.

大别山北缘不同生活型木本植物叶功能性状及其耦合关系

【目的】探讨大别山北缘不同生活型木本植物叶功能性状变异,研究叶功能性状指标之间的耦合关系,为解析植物对气候变化的响应机制提供依据。【方法】采集大别山北缘连康山国家级自然保护区30种常见的不同生活型木本植物(常绿、落叶、乔木、灌木)的叶片,测定其叶绿素含量(以SPAD表示)、干鲜比、氮质量分数、磷质量分数、钾质量分数、氮磷比、叶面积、比叶面积共8个叶功能性状指标,分析了不同生活型木本植物叶功能性状及其相关性。【结果】不同生活型木本植物叶功能性状指标具有一定的差异性,其中常绿和落叶树种的叶片SPAD、氮质量分数、磷质量分数、钾质量分数、叶面积和比叶面积之间差异显著(P<0.05);乔木与灌木树种的叶面积、氮质量分数、磷质量分数、钾质量分数、干鲜比与氮磷比差异显著(P<0.05)。部分叶功能性状指标之间存在一定的相关性,叶片氮、磷质量分数与钾质量分数呈显著正相关(P<0.05),叶片氮质量分数与磷质量分数呈显著正相关(P<0.05);叶面积与叶片氮、磷、钾质量分数呈显著正相关(P<0.05),比叶面积与叶片氮、磷、钾质量分数呈显著正相关(P<0.05)。主成分分析结果表明:叶片磷、氮、钾质量分数是较为重要的叶功能性状指示指标,可将30种木本植物归为4个功能组,且第1和第2主成分对植物叶功能性状变异的贡献率分别为55.2%和12.8%。【结论】叶功能性状指标在不同生活型木本植物之间存在一定差异,其中叶磷、氮、钾质量分数是指示性指标,且与叶面积显著正相关。图4表1参28.