Population structure,age and growth rates of conifer species and their relation to environmental variables at Malam Jabba,Swat District,Pakistan

This study focuses on age,growth rate and diameter distribution of pine forests in the Malam Jabba area,Swat District,Pakistan.Wood core samples were taken from twenty stands.Picea smithiana was the oldest at 234 years with a 112-cm diameter.Abies pindrow was 125 years with an 80-cm diameter while the oldest Pinus wallichiana was 122 years with 75-cm diameter.The fastest overall growth rate of 1.5±0.1 year/cm was for P.wallichiana on a west-facing aspect,while the slowest 5.8±2.6 year/cm growth was P.smithiana on an eastfacing exposure.P.wallichiana and A.pindrow exhibited marked differences in growth rates over a 5-year period.The highest growth was by P.wallichiana from 1966 to 2006.A.pindrow showed less growth over the same years,such pattern simultaneously reverse from 1911 to 1965.The relationship between diameter and age,diameter and growth rate and age and growth rate were correlated.P.wallichiana and A.pindrow ages were correlated with diameter and growth rates.P.smithiana age was positive correlated with diameter.Generally,topographic and edaphic factors did not show significant correlations with growth rates,although some appreciable correlations were recorded.The growth of P.wallichiana was correlated with elevation while A.pindrow was correlated with maximum water retaining capacity.Diameter and age produced uneven size classes and many size gaps,which could be the result of anthropogenic disturbances.

Forest biomass carbon storage from multiple inventories over the past 30 years in Gansu Province, China: implications from the age structure of major forest types

We used the forest inventory data of Gansu Province, China to quantify carbon storage and carbon density changes by regional forest cover and by typical forest types in 1979-2006. Total forest area increased from 1.77 x 106 ha in 1979 to 2.32 x 106 ha in 2006, and the forest carbon storage, estimated by the continuous biomass expansion factor method, increased from 83.14 to 100.66 Tg, equivalent to a carbon accumulation rate of 0.0071 Tg per year during the period. Mean carbon densities were 44.83-48.50 t ha-1 and the values decreased slightly over the time period. Natural forests generated greater car- bon storage and density than did plantations. By regression analysis, forest stand age was an important parameter incarbon density studies. We developed various regression equations between carbon density and stand age for major types of natural forests and plantations in the region. Our results can be used for proper selection of re-forestation species and efficient management of young and middle-aged forests, offering great potential for future carbon sequestra- tion, especially in arid and semi-arid regions.

Factors Associated with District Tsunami Victim Rate by Age Class in Ishinomaki City： 2011 Great East Japan Earthquake and Tsunami

This paper analyzes the factors associated with the district tsunami victim rate in 2011 by age class/group in Ishinomaki city, Japan. Sea or river visibility and evaluation potential from the temporary facilities to higher hills are factors associated with victim rate in addition to tsunami depth. In addition, victim rate, especially for elder people, is low in the districts that have higher aging rate. The characteristics of these districts are： （1） many people are engaged in fishery related jobs and know each other and （2） higher hills are behind the beach.

Genetic Variation and Differentiation of Larix decidua Populations in Swiss Alps

Genetic diversity within and among six subpopulations of Larix decidua Mill. from two altitudinal transects of Swiss Alps was investigated using 6 enzyme systems coding for 8 loci. Globally, the mean proportion of polymorphic loci was 22.9%, the average number of alleles per locus was 1.3, and the mean expected heterozygosity was 0.095. Only 5.8% of the genetic variation resided among populations. The mean genetic distance was 0.006. Several significant differences of gene frequencies were found between different age classes. Positive values of the species mean fixation index observed in this study suggested a considerable deficit of heterozygotes in the populations of L. decidua of Swiss Alps. At one of the sites (Arpette), the highest subpopulation in elevation gave the lowest level of genetic diversity (as evidenced by the lowest proportion of polymorphic loci and the lowest mean expected heterozygosity) and the largest value of genetic distance when compared to other subpopulations. The genetic differences between the highest subpopulation and the other ones suggest that the founder effect may be an important factor influencing genetic differentiation of L. decidua populations at Arpette transect.

Comparison of the water-crossing behavior of Western Marsh Harriers(Circus aeruginosus) and European Honey Buzzards(Pernis apivorus) during autumn migration

We investigated the water-crossing behavior of Western Marsh Harriers(Circus aeruginosus),mostly adults,and European Honey Buzzards(Pernis apivorus),mostly juveniles,in relation to wind conditions,time of the day,flocking and age classes,at a watchsite in central Italy during the autumn migration en route to Africa.Although European Honey Buzzards are less suited than Western Marsh Harriers to undertake long sea crossings,they were more inclined to leave the coast when migrating in flocks.Few birds of both species chose to fly along the coast.Western Marsh Harriers,such as European Honey Buzzards migrating alone,undertook the water crossing rather than stopping migration(birds roosting at the site or flying back inland) during the absence of wind and vice versa during head winds.Conversely,European Honey Buzzards migrating in flocks were not affected in their decision(crossing or stopping migration) by wind direction.Both species undertook the water crossing rather than stopping migration during mornings and vice versa during afternoons.Finally,in both species,adults and juveniles showed the same behavior in front of a water barrier.This result was expected in the case of the Western Marsh Harrier but not from the European Honey Buzzard since,in this species,the water-crossing tendency is age dependent with adults avoiding sea crossings.Our study confirms that flocking significantly affects the water-crossing behavior of European Honey Buzzards during migration.Moreover,in this species,inexperience of juveniles and presumably younger adults,about the high energetic costs of long powered flight and about the existence of shorter routes over water,might explain the strong water-crossing tendency shown by migrants independently from their age.

Spatial distribution pattern of Picea schrenkiana population in the Middle Tianshan Mountains and the relationship with topographic attributes

The spatial distribution of plant populations is an important feature of population structure and it de- termines the population＇s ecological preferences, biological characteristics and relationships with environmental factors. The point pattern analysis method was adopted to study the distribution pattern of Picea schrenkiana indi- viduals of different size classes and the correlations between two size classes as well as the impact of topog- raphical attributes on the population distribution. With increasing diameter at breast height, the plant density of the P. schrenkiana population showed a declining trend. Old trees showed a random distribution at a small spatial scale （0-12 m）, whereas saplings, small trees and big trees all had an aggregated distribution at all scales. With the increase of tree age, the scales at which maximal aggregation occurred gradually increased and the aggregation strength decreased. At a small scale （0-16 m）, all size classes showed a negative correlation and the larger the difference between tree size, the more significant the negative correlation. The number of medium, big and old trees had a significantly positive correlation with elevations, whereas the number of saplings and small trees was not significantly correlated with elevations. The numbers of saplings, small and medium trees showed a significant positive correlation with slope gradient, whereas the number of big trees was not significantly correlated, and the number of old trees was negatively correlated with gradient. With the exception of old trees, saplings, small, me- dium and big trees showed negative correlations with convexity index. The study provides a theoretical basis for the conservation, rehabilitation and sustainable management of forest ecosystems in the Tianshan Mountains.

Activity budgets in a group of Sichuan snub-nosed monkeys in Shennongjia Nature Reserve,China

We examined time allocation by Sichuan snub-nosed monkeys Rhinopithecus roxellana in the montane, temperate and highly seasonal forests of the Shennongjia Nature Reserve in China, in order to improve our understanding of the ecological and social influences on monkey behavior. We collected data on activity budgets in relation to food availability in a group of monkeys from July 2003 to September 2004 （except February 2004）, using instantaneous scan samples. The monkeys spent 36.21% of daytime moving （ n = 21, 269 records）, 24.39% foraging, 9.33% inactive, 11,7% performing social behaviors, and 18.38% performing other behaviors. There were seasonal and annual variations in the proportions of time devoted to some activities. These variations were largely attributed to seasonal or annual fluctuations in the distribution and quality of food resources. Adult males spent more time moving, searching, and displaying other behaviors, but less time feeding and performing social behaviors than adult females. Adults performed other behaviors more often and moving less often than juveniles. These differences in activity budgets may reflect fundamental differences in reproductive biology, parental investment, and development among the different age/sex classes [ Current Zoology 55 （3） ： 173 - 179, 2009] .

Ontogenetic transition in leaf traits: a new cost associated with the increase in leaf longevity

Aims Recent work has identified a worldwide‘economics’spectrum of correlated leaf traits that mainly reflects the compromises between maximizing leaf longevity and short-term productivity.However,during the early stages of tree growth different species tend to exhibit a common strategy,because competition for soil water and nutrients forces the maximization of short-term productivity owing to the need for rapid growth during the most vulnerable part of the tree’s life cycle.Accordingly,our aim here was to compare the variations that occur during ontogeny in the different leaf traits(morphology and leaf chemical composition)of several coexisting Mediterranean woody species differing in their leaf life spans and to test our hypothesis that tree species with a long leaf life span should exhibit larger shifts in leaf characteristics along ontogeny.Methods Six Mediterranean tree species differing in leaf life span,selected from three plots located in central-western Spain,were studied during three growth stages:seedlings,juveniles and mature trees.Leaf life span,leaf morphology(leaf area,dry weight,thickness and mass per unit area)and chemical composition(N and fibre con-centrations)were measured in all six species.The magnitude of the ontogenetic changes in the different traits was estimated and related to the mean leaf longevity of the different species.Important Findings Along ontogeny,strong changes were observed in all variables analysed.The early growth stages showed lower leaf thickness,leaf thickness and mass per unit area and N,cellulose and hemi-cellulose concentrations than mature trees,but a higher lignin content.However,these changes were especially marked in species with a longer leaf life span at maturity.Interspecific dif-ferences in leaf life span,leaf morphology and chemical com-position were stronger at the mature stage than at the seedling stage.We conclude that greater plasticity and more intense strat-egy shifts along ontogeny are necessarily associated with long leaf life span.Our results thus provide a new aspect that should be incorporated into the analysis of the costs and benefits associ-ated with the different strategies related to leaf persistence dis-played by the different species.Accordingly,the intensity of the alterations in leaf traits among different growth stages should be added to the suite of traits that change along the leaf economics spectrum.