Soil carbon dioxide fluxes of a typical broad-leaved/Korean pine mixed forest in Changbai Mountain, China

The forest ecosystem plays an important role in the global carbon cycling. A study was conducted to evaluate soil CO2 flux and its seasonal and diurnal variation with the air and soil temperatures by using static closed chamber technique in a typical broad-leaved/Korean pine mixed forest area on the northern slope of Changbai Mountain, Jilin Province, China. The experiment was carried out through the day and night in the growing season (from June to September) in situ and sample gas was analyzed by a gas chromatograph. Results showed that the forest floor was a large net source of carbon, and soil CO2 fluxes had an obvi-ous law of seasonal and diel variation. The soil CO2 flux of broad-leaved/Korean pine mixed forest was in the range of 0.302.42 mmol穖-2穝-1 with the mean value of 0.98 mmol穖-2穝-1. An examination on the seasonal pattern of soil CO2 emission suggested that the variability in soil CO2 flux could be correlated with variations in soil temperature, and the maximum of mean CO2 flux occurred in July ((1.27±23%) mmol穖-2穝-1) and the minimum was in September ((0.50±28%) mmol穖-2穝-1). The fluctuations in diel soil CO2 flux were also correlated with changes in soil temperature; however, there existed a factor for a time lag. Soil CO2 flux from the forest floor was strongly related to soil temperature and had the highest correlation with temperature at 6-cm depth of soil. Q10 values based on air temperature and soil temperature of different soil depths were at the ranges of 2.09–3.40.

Factors controlling N_2O and CH_4 fluxes in mixed broad-leaved/Korean pine forest of Changbai Mountain, China

Using the closed chamber technique, the in situ measurements of N2O and CH4 fluxes was conducted in a broad-leaved Korean pine mixed forest ecosystem in Changbai Mountain, China. from June 1994 to october 1995. The relationships between fluxes (N2O and CH4) and some major environmental factors (temperature, soil water content and soil availabIe nitrogen) were studied. A significant positive correlation between Nzo emission and air/soil temperature was observed, but no significant correIation was found between N2O emission and soil water content (SWC). This result showed that temperature was an important controlling factor of N2O flux. There was a significant correlation between CH4 uptake and SWC, but no significant correlation was found between CH4 uptake and temperature. This suggested SWC was an important factor controlling CH4 uptake. The very significant negative correlation between logarithmic N2O flux and soil nitrate concentration, significant negative correlation between CH4 flux and soil ammonium content were also found.

Dynamics of nitrogen nutrition of coexisting dominant trees in mixed broad-leaved/Korean pine forest

Chemical analysis of ammonium, nitrate and total nitrogen in tree leaves and roots and anin-vivo bioassay for nitrate reductase activity (NRA) were used to monitor the seasonal variations in nitrogen assimilation among four coexisting dominant tree species, includingPinus koraiensis, Tilia amurensis, Fraxinus mandshurica andAcer mono, in a virgin mixed broad-leaved/Korean pine (Pinus koraiensis) forest. The soil study included individual horizons of L+F (0–5 cm), Ah (5–11 cm) and Aw (11–25 cm). All four species had nitrate and ammonium in their roots and leaves, and also NRA in leaves. This indicated that these coexisting species were adapted to ammonium + nitrate nutrition. A negative correlation existed between nitrate use and ammonium use. Ammonium concentration was higher than that of nitrate in tree leaves and roots, and also in soils, which indicated climax woody species had a relative preference for ammonium nutrition. There was a positive relationship between tree nitrogen nutrition use and soil nitrogen nutrient supply. Utilization of ammonium and nitrate as well as the seasonal patterns differed significantly between the species. Peaks of ammonium, nitrate, NRA and total nitrogen in one species were therefore not necessarily synchronous with peaks in other species, and which indicated a species-specific seasonal use of nitrogen. The species-specific temporal differentiation in nitrogen use might reduce the competition between co-existing species and may be an important mechanism promoting stability of virgin mixed broad-leaved//Korean pine forest.

NITROGEN NUTRIENT MECHANISM IN SECONDARY SUCCESSION PROCESS OF THE MIXED BROAD-LEAVED/ KOREAN PINE (PINUS KORAIENSIS) FOREST

Chemical and biochemical analysis methods were used to monitor the vedations of nitrogen nutrient among the dominance trees species in secondary succession process of the mixed broad - leaved/Korean pine forest on Changbai Mountains, Northeast China. Amounts of total nitrogen, anunonium and NRA in soils of virgin broad-leaved/Korean pine forest which is in climax were higher than those of secondary birch forests those are in succession Stage. The amount of nitrate was in the other hand. In climax, dominance trees species are tolerant mesophytic trees such as Pinus Koraiensis, Tilia amurensis, Acer mono and also Fraxinus mandshurica, they are all ammonium + nitrate adapted species, but they show a preference for the anunonium rather than those of the pioneer trees species in secondary birch forest, such as Populus davidiava and Betula platyphylla. Because they have more ammonium in their leaves and roots, especially Pinus koraiensis. Populus davidvana and Betula plaaphlla are intolerant trees, amounts of nitrate and total nitrogen is higher in their leaves and roots and also NRA in their leaves, so they preference for the nitrate rather than the others.In secondary birch forest, the regeneration trees species adapt their nitroggn nutrient to the variation of nitrogen nutrient situation in soil, finally they could survival well and the secondary birch forest would succession to climax. In climax, dominance trees species adapt their Nitrogen nutrient to the situation in soil and there are not strong competition in nitrogen nutrient among them, so they can coexist well and keep the climax as stable vegetation.

Long-term changes in radial growth of seven tree species in the mixed broadleaf-Korean pine forest in Northeast China:Are deciduous trees favored by climate change?

The role of the temperate mixed broadleaf-Korean pine forest(BKF)in global biogeochemical cycles will depend on how the tree species community responds to climate;however,species-specific responses and vulner-abilities of common trees in BKF to extreme climates are poorly understood.Here we used dendrochronological meth-ods to assess radial growth of seven main tree species(Pinus koraiensis,Picea jezoensis,Abies nephrolepis,Fraxinus mandshurica,Phellodendron amurense,Quercus mongolica,and Ulmus davidiana)in an old-growth BKF in response to climate changes in the Xiaoxing’an Mountains and to improve predictions of changes in the tree species compo-sition.Temperature in most months and winter precipita-tion significantly negatively affected growth of P.jezoensis and A.nephrolepis,but positively impacted growth of P.koraiensis and the broadleaf species,especially F.mandshu-rica and U.davidiana.Precipitation and relative humidity in June significantly positively impacted the growth of most tree species.The positive effect of the temperature during the previous non-growing season(PNG)on growth of F.mandshurica and Q.mongolica strengthened significantly with rapid warming around 1981,while the impact of PNG temperature on the growth of P.jezoensis and A.nephrolepis changed from significantly negative to weakly negative or positive at this time.The negative response of radial growth of P.jezoensis and A.nephrolepis to precipitation during the growing season gradually weakened,and the negative response to PNG precipitation was enhanced.Among the studied species,P.koraiensis was the most resistant to drought,and U.davidiana recovered the best after extreme drought.Ulmus davidiana,P.jezoensis and A.nephrolepis were more resistant to extreme cold than the other species.Climate warming generally exacerbated the opposite growth patterns of conifer(decline)and broadleaf(increase)spe-cies.Deciduous broadleaf tree species in the old-growth BKF probably will gradually become dominant as warming continues.Species-specific growth-climate relationships should be considered in future models of biogeochemical cycles and in forestry management practices.

Recovery of species diversity after disturbance of broad-leaved Korean pine mixed forest in Changbai Mountain

Recovery of species diversity after catastrophic disturbance was influenced by a few factors, such as intensity of disturbance, availability of propagules, and the environmental conditions, In this paper, we examined pattems of species development after nearly 60a succession in bumed broad-leaved Korean pine mixed forest on northem slope of ChangbaiMountain. We aasessed the recovery of plant species diversity in 3 types of forests under the contition with gradient of soil moisture. Results revealed that recovery of plant species (liversity varied greatly under different environmental conditions.Species richness of secondary forests greatly related to the site condition. Secondary birch forest on mesic site had the greatest number of plant species and the following was poplar-birch forest and larch-birch forest.Most of characteristic taxa couldbe found in birch forest after 60a succession. For larch-birch forest on hydra site, most of climax species were still not able to invade, so it had the lowest species diversity.

Species composition and minimum sampling area of a riparian mixed broadleaved-Korean pine forest in Changbai Mountain Nature Reserve

Riparian areas are unique although often small component of the overall watershed landscape. The structure of riparian forests along Erdaobai River on the north slope of Changbai Mountain were investigated by using field data collected from eight sampling transects perpendicular to the Erdaobai River channel. Two kinds of species-area saturation curves were used to examine the relationship between species number and minimum sampling area. The results showed that riparian gymnosperms accounted for a high proportion of all gymnosperms in the Changbai Mountain Nature Reserve while riparian ferns and angiosperms accounted for a relatively low proportion. The average minimum sampling areas of riparian forest that included 60%, 80%, and 90% of the community species pool were about 85, 185, and 328 m 2 , respectively; while those for nonriparian forest were about 275, 390, and 514 m 2 , correspondingly.

Age structure of Tilia broad-leaved Korean pine forest on northern slope of Changbai Mountain

Based on a vast of field investigation on stamps in Tilia broad-leaved Korean pine forest on northem slope of Changbai Mountain, age structure of some major species were studied in this paper. The results showed that Korean pine population was composed of grouped patches with different ages. There were not strict intervals among the dominated generations,and the curve of age structure often had two or more peaks. The distribution of broad-leaved species in natural Korean pine forest was grouped or scattered, and age distribution was also uneven-aged. There existed close relation between quantity of broad-leaved species and Korean pine. So, it shaped multi-storied and uneven-aged mixed forest. The model of age structure and growth demonstrated their passive correlation, but growth became slow when woods had reached old age.

A succession and silviculture model for the broad-leaved Korean pine forests of Changbai Mountain Area

A succession and silviculture model (ZELIG.CBA) for broad-leaved Korean pine forest of Changbai Moutain Area was developed based on the framework of ZELIG model and characteristics of Broad-leaved Korean pine forests of Changbai area. ZELIG.CBA model consists four sub-models: growth model simulating annual increment of individual tree in forest, regeneration model simulating annual establishment of different tree species, mortality model simulating annual agerelated and stress-related morality; and silviculture model simulating the forest response to different silviculture scenario. The validation of the ZELIG.CBA showed that the accuracy of the model for the forest growth was more than 95%. The succession from clear cutting site simulating showed that the ZELIG.CBA model was stable for long term simulation. And selective cutting experiment showed that the optimal scenario for broad-leaved Korean pine forests was removal volume 30% combining with 30a rotation.

Canopy gap characteristics and its influence on the regeneration of broad-leaved Korean pine forests in Changbai Mountain

With the concem of regcneration, characteristics of canopy gaps in broad-leaved Korean pine forest were studied.The areas of actual gap, expanded gap and maximum gap were analyzed respectively. The species composition, number, origin,decayed class and sizes of gap makers were studied comprehensively. The comparative studies of regencration inside and outside of canopy gap showed that the density of regeneration inside canopy gaps was 30% higher than that outside canopy gaps. The specific species regeneration response to canopy gap varied greatly. Pinus koraiensis, Picea jezoensis Frarinlis nla)ldshurica, JItghI)ls )nalJholu-i`a and Acer mono generally did not response to canopy gap disturbance. The Regeneration Importantce Valuc (RIV) of Abies nephrolepis, Ulmus Japonica increased with canopy gap disturbance. RIV of Tilia amurensis, Acer madshurica and Ulmus laciniata decreased with canopy gap disturbance. Canopy gap disturbance was not strong enough to result in the regeneration of some rpecies of shade intolerance such as Larix olgensis, Betula platyphylla.

Meteorological control on CO_2 flux above broad-leaved Korean pine mixed forest in Changbai Mountains

The impacts of temperature, photosynthetic active radiation (PAR) and vapor pressure deficit (VPD) on CO2 flux above broad-leaved Korean pine mixed forest in the Changbai Mountains were studied based on eddy covariance and meteorological factors measurements.The results showed that, daytime CO2 flux was mainly controlled by PAR and they fit Michaelis-Menten equation. Meanwhile VPD also had an influence on the daytime flux. Drier air reduced the CO2 assimilation of the ecosystem, the drier the air, the more the reduction of the assimilation. And the forest was more sensitive to VPD in June than that in July and August. The respiration of the ecosystem was mainly controlled by soil temperature and they fit exponential equation. It was found that this relationship was also correlated with seasons; respiration from April to July was higher than that from August to November under the same temperature. Daily net carbon exchange of the ecosystem and the daily mean air temperature fit exponential equation. It was also found that seasonal trend of net carbon exchange was the result of comprehensive impacts of temperature and PAR and so on. These resulted in the biggest CO2 uptake in June and those in July and August were next. Annual carbon uptake of the forest ecosystem in 2003 was -184 gC. m-2.

Modeling diameter distribution of the broadleaved-Korean pine mixed forest on Changbai Mountains of China

The broadleaved-Korean pine mixed forest is a native vegetation in the Changbai Mountains, northeast China. The probability density functions including the normal, negative exponential, Weibull and finite mixture distribution, were used to describe the diameter distributions of the species groups and entire forest stand. There is a strong correlation between parameters and mean DBH except the shape parameters in the mixture distribution. The diameter classes of species and entire forest stand showed not negative exponential but normal and "S" distribution. The mixture function was better than normal and Weibull to describe the model distribution. The location parameter had an effect on the estimated frequency in the first diameter class, when the estimated location parameter was bigger than the lower limit of the first diameter class.

长白山主要林分物种多度分布与性状分布的关系

【目的】探究长白山地区群落物种的多度分布格局及其驱动过程,以及多度分布与性状分布之间的关系,为当地森林经营与修复策略的制定提供理论依据。【方法】以长白山不同林分类型的3块5.2 hm2(次生山杨白桦林、次生针阔混交林、原始红松紫椴林)和1块5 hm2原始阔叶红松林固定监测样地为研究对象,通过采集样地内木本个体的6个关键功能性状(叶面积、比叶面积、叶片厚度、最大树高、叶片氮含量、叶片磷含量),分析样地的物种多度分布格局、功能性状分布格局以及二者之间的关系。【结果】不同林分类型中通过统计检验的模型种类各异,但所有最优模型均为统计模型。在性状分布格局中,最大树高、比叶面积和叶磷含量在4块样地均呈现正态分布。然而,功能性状分布格局转化得到的物种分布格局模型显示,尽管部分模型通过了统计检验,但与多度分布结果拟合度不高。【结论】在长白山地区,物种多样性的形成并非主要由随机过程驱动,而是受到竞争作用的显著影响,这决定了演替过程中群落物种组成的变化情况。虽然功能性状分布与物种多度分布之间存在一定的联系,但直接通过群落水平性状分布格局推断物种多度的方法尚不准确。

长白山阔叶红松林林层群落结构与生产力的关系

【目的】探究长白山阔叶红松林不同林层群落结构与林分生产力的关系,分析林分生产力的影响因素,为温带森林的恢复与可持续经营提供科学依据。【方法】基于长白山自然保护区阔叶红松林40 hm^(2)固定监测样地2年(2014和2019年)植被调查数据,以样地内胸径(DBH)≥5 cm的木本植物为对象,根据胸径将阔叶红松林群落划分为优势木层、亚优势木层、中等木层和被压木层,运用R4.0.3软件计算各林层物种多样性、结构多样性和林分密度,与生产力进行相关性分析并构建结构方程模型,探讨不同林层物种多样性、结构多样性和林分密度对生产力的影响。【结果】1)林分密度与生产力显著正相关(P<0.05),且相关关系随林层上升而下降。2)Pearson相关性分析结果显示,生产力与代表物种组成α多样性的香农威纳指数显著正相关(P<0.05),且随林层上升相关关系减弱;反之,代表物种组成β多样性的香农熵指数与生产力存在显著负相关(P<0.05),但该相关关系在中等木层和亚优势木层不显著。此外,物种均匀度指数与生产力显著负相关(P<0.05),相关系数随林层上升先增大后减小;胸高断面积基尼系数和胸径均匀度指数与生产力显著负相关(P<0.05),且该相关关系随林层上移而减弱,在优势木层与生产力无显著相关性。胸径香农威纳指数与生产力呈显著正相关(P<0.05),且随林层上移相关关系降低,相关系数由0.44降至0.31。3)不同林层林分密度对生产力均有显著影响(P<0.001),且随林层上升其作用强度逐渐降低,路径系数由0.59降至0.27;物种多样性在被压木层和中等木层对生产力有显著正向作用(P<0.05),在优势木层和亚优势木层对生产力无显著影响;结构多样性在被压木层和中等木层通过作用于物种多样性和林分密度间接影响林分生产力,在优势木层和亚优势木层则直接作用于林分生产力。【结论】各林层林分密度对生产力均有显著影响(P<0.05),且随林层上升对生产力的作用强度逐渐降低,群落物种多样性和结构多样性对生产力的作用方向和强弱在不同林层表现各异。深入探讨不同林层物种多样性和结构多样性以及其如何影响森林生产力,在现代森林经营管理中尤为重要。

吉林蛟河阔叶红松林主要树种叶片养分含量及氮磷再吸收特征

以吉林蛟河阔叶红松林常绿乔木(红松)和3种落叶乔木(蒙古栎、紫椴和色木槭)为研究对象,采集4种乔木成熟叶与衰老叶,测量其碳(C)、氮(N)、磷(P)含量,并计算其化学计量比和养分再吸收效率,分析不同生活型乔木叶片养分含量、化学计量比和养分再吸收效率的差异,研究不同生活型乔木养分保存机制。结果表明:(1)落叶乔木叶片N、P含量显著高于红松,衰老叶N含量差异不显著。(2)红松成熟叶C∶N、C∶P显著高于落叶乔木,红松衰老叶C∶N显著高于落叶乔木,不同生活型乔木叶片N∶P没有显著差异。(3)落叶乔木N再吸收效率(NRE)和P再吸收效率(PRE)显著高于红松;乔木叶片NRE/PRE比值均大于1。(4)乔木成熟叶N、P含量均与NRE和PRE呈显著正相关关系,衰老叶C∶N和C∶P均与NRE和PRE呈显著负相关关系。总之,吉林蛟河阔叶红松林4种乔木成熟叶养分含量会影响其养分再吸收效率,衰老叶C∶N和C∶P可以反映其养分再吸收效率,4种乔木均更倾向于从衰老叶片中吸收N。

胜山阔叶红松林粗木质残体空间分布格局

为探究胜山阔叶红松(Pinus koraiensis)林粗木质残体(Coarse woody debris,CWD)的形成过程及机制,采用点格局分析法对10.4 hm^(2)大型样地内不同分类的CWD的空间格局、空间关联性进行了研究。结果表明:(1)总CWD在0—7 m尺度聚集分布,7—50 m尺度随机分布。(2)腐烂等级5级CWD、倒木、伐根在所有尺度随机分布;其他CWD随尺度增大由聚集分布转为随机分布,但格局尺度不同(在2—10 m之间)。(3)大、中径级与小径级CWD在0—4 m、0—6 m尺度正相关;高腐烂等级与低腐烂等级CWD在0—5 m尺度以内正相关;枯立木与倒木在0—4 m尺度正相关;落叶松(Larix gmelinii)CWD与小径级、腐烂等级2级的红皮云杉(Picea koraiensis)CWD在0—3 m尺度正相关,与中径级、腐烂等级1、2级在13—16 m尺度负相关。胜山阔叶红松林CWD的形成是由树种特性、小尺度的个体竞争、大尺度的自然衰老、外界干扰、生境异质性等共同决定的;大、中径级对小径级CWD、先形成对后形成CWD、枯立木对倒木具有一定的正向影响,落叶松对红皮云杉CWD在小尺度有正向影响,而在稍大尺度有负面影响;在一定程度上揭示了该林型CWD的形成过程和机制。

2011-2020年长白山阔叶红松林凋落物回收量和现存量动态数据集

长白山阔叶红松林是我国目前温带面积最大、保护最为完整的森林生态系统之一。森林凋落物作为森林地上净生产量回归土壤的主要方式,是森林生态系统养分归还的重要途径。森林凋落物研究对于理解森林碳循环以及预测其对气候变化的响应都有着极为重要的意义。作为中国生态系统研究网络(Chinese Ecosystem Research Network,CERN)和国家野外科学观测研究站网络(National Ecosystem Research Network of China,CNERN)成员,长白山森林生态系统定位研究站按照CERN的统一规范,在长白山阔叶红松林野外永久样地安装10个凋落物框,长期对其进行凋落物回收量监测。本数据集通过整理和统计,汇总了2011-2020年长白山阔叶红松林凋落物各组分(枯枝、枯叶、落果(花)、树皮、苔藓地衣及杂物的干重)回收量月动态及现存量年动态数据,并包含了相关的数据集构建过程信息,数据集可为森林生态和土壤研究提供基础。

台风对小兴安岭阔叶红松林的影响

台风对森林造成强烈干扰,改变森林生态系统的结构与功能。分析了2020年台风对小兴安岭地区阔叶红松(Pinus koraiensis)林中不同径级、功能群(耐阴性、生活型、针阔叶)树木破坏程度的影响和地形(海拔、坡向、坡度、凹凸度)在其中的作用。结果表明:台风对不同径级以及不同功能群树种的受损数量的影响具有显著差异,具体表现为径级Ⅰ(胸径<10 cm)>径级Ⅱ(10 cm≤胸径<50 cm)>径级Ⅲ(胸径≥50 cm),耐阴性强>耐阴性弱、乔木>灌木、阔叶树种>针叶树种;而其受损率则表现为径级Ⅱ>径级Ⅲ>径级Ⅰ,耐阴性弱>耐阴性强、乔木>灌木、针叶树种>阔叶树种;损坏程度很大程度上取决于坡向与海拔位置,处于阴坡与低海拔的受损木数量较高,阔叶树种、灌木与耐阴性强的受损木表现出在阳坡与海拔较高位置出现的可能性更大,针叶树种、乔木与耐阴性弱的受损木表现出在阴坡与海拔较低位置出现的可能性更大,阴坡和较低海拔处拔根倒的受损数量显著高于干基折和干中折的受损数量;地形对受损木的风倒方向具有显著影响。总体而言,台风对森林的破坏受到多种环境因素和生物因素共同作用的影响,处于不同地形的不同径级、功能群的树木,其对台风的抵抗能力有所不同,且大径级树木、耐阴性弱的树种、乔木和针叶树种对台风的抵抗能力相对较弱,迎风坡与阴坡的树木对台风的抵抗能力相对较弱,这为理解极端天气事件对森林动态的影响以及恢复提供理论依据与参考。

阔叶红松林皆伐后不同恢复方式下土壤有机碳形态与矿化速率变化

土壤有机碳矿化对森林恢复方式有极强的敏感性,为阐明不同恢复方式下土壤有机碳矿化速率及其与碳形态之间的关系,2023年6月选择小兴安岭地区阔叶红松林皆伐迹地区域内2种不同恢复方式下形成的2种林分(天然恢复形成的天然枫桦次生林,人工恢复形成的红松人工林)的林地土壤作为研究对象,以未受干扰的原始阔叶红松林的林地土壤作为对照(PK),采用野外样地设置与调查、土样采集与室内培养及测定相结合的方法,分析有机碳矿化速率对不同恢复方式的响应,并结合土壤碳形态的分布,探究不同恢复方式下有机碳矿化速率与土壤碳形态之间的关系。研究结果表明,1)土壤有机碳矿化速率从第3天开始随着培养时间的延长呈现出不断下降直至趋于平稳的趋势,且矿化速率随着土层的加深逐渐减小;2)原始阔叶红松林(PK)的矿化速率最高、天然枫桦次生林(BC)次之、红松人工林(PM)最低;3)2种恢复方式下土壤的总有机碳、矿物结合态有机碳与非活性有机碳由大到小依次为PK、BC、PM;颗粒有机碳含量由大到小依次为BC、PM、PK;4)培养过程中土壤总有机碳、颗粒有机碳、易氧化有机碳含量与有机碳矿化速率均呈现出一致减小的趋势,且与有机碳矿化量呈显著负相关性,其中易氧化有机碳对土壤有机碳矿化率的影响最大,相关系数为0.622。矿物结合态有机碳和非活性有机碳与土壤有机碳矿化量无显著相关性。综上,相较于红松人工林,天然枫桦次生林增强了土壤有机碳矿化速率,并且在2种恢复方式中,活性有机碳主导着土壤有机碳矿化速率。

阔叶红松林中黑啄木鸟的取食生境偏好

2020年冬季—2022年冬季,采用样线法和样方法对黑龙江省凉水国家级自然保护区中黑啄木鸟(Dryocopus martius)取食生境偏好进行调查研究,结果表明:在取食微生境尺度中,黑啄木鸟取食树树种、取食树状态与对照样方具有显著差异(p<0.05),偏好在冷杉(Abies fabri,47.71%)和半枯立木(73.86%)的树木主干(86.93%)中上部(43.14%)、上部(32.03%)取食,且偏好粗壮、高大的树木;在取食样方尺度中,黑啄木鸟偏好在有林窗(92.16%)的天然林(96.73%)中取食,偏好郁闭度较低、优势乔木均高较低、优势乔木平均胸径较大、样方中活立木较少、枯立木、倒木较多和距人为干扰更近的生境中取食,与对照样方呈显著差异(p<0.05)。资源选择函数结果表明,影响黑啄木鸟取食生境偏好的关键因子是取食树树种,次关键因子是枯立木数量、优势乔木均高、取食树高度和倒木数量。