Species selection in hardwoods research: variations in baseline physiological responses of select temperate hardwood tree species

Drought periods are becoming more extreme worldwide and the ability of plants to contribute towards atmospheric flux is being compromised. Properly functioning stomata provide an exit for water that has been absorbed by the roots, funneled into various cell parts, and eventually released into the atmosphere via transpiration. By observing the effects that weather conditions such as climate change may have on stomatal density, distribution, and functioning, it may be possible to elucidate a portion of the mechanisms trees use to survive longer periods of water stress. This study analyzed stomatal density （SD）, stomatal conductance （gs ）, CO2 assimilation （A）, instantaneous water-use efficiency （WUEi ）, and transpiration （E） rates in six native tree species in the Midwestern USA and showed that trees within the same ecotype followed similar trends, but that trees within the same family did not when exposed to identical greenhouse conditions. Naturally drought tolerant tree species demonstrated lower g s and higher WUEi , while intolerant species had higher SD. This study showed negative or no correlation between SD and g s , A, E, and WUEi and positive correlations between E and A and gs and E.

Simulation of Steel Reinforcement on the Nonlinear Behaviour of Slender Glulam Beam Columns by Using the Newton-Raphson Method

The current theory in NF EN 1995-1-1/NA of Eurocode 5, which is based on maximum deflection, has been investigated on softwoods. Therefore, this theory is not adapted for slender glulam beam columns made of tropical hardwood species from the Congo Basin. This maximum deflection is caused by a set of loads applied to the structure. However, Eurocode 5 doesn’t provide how to predict this deflection in case of long-term load for such structures. This can be done by studying load-displacement (P-Δ) behaviour of these structures while taking into account second order effects. To reach this goal, a nonlinear analysis has been performed on a three-dimensional beam column embedded on both ends. Since conducting experimental investigations on large span structural products is time-consuming and expensive especially in developing countries, a numerical model has been implemented using the Newton-Raphson method to predict load-displacement (P-Δ) curve on a slender glulam beam column made of tropical hardwood species. On one hand, the beam has been analyzed without wood connection. On the other hand, the beam has been analyzed with a bolted wood connection and a slotted-in steel plate. The load cases considered include self-weight and a uniformly applied long-term load. Combinations of serviceability limit states (SLS) and ultimate limit states (ULS) have also been considered, among other factors. A finite-element software RFEM 5 has been used to implement the model. The results showed that the use of steel can reduce displacement by 20.96%. Additionally, compared to the maximum deflection provided by Eurocode 5 for softwoods, hardwoods can exhibit an increasing rate of 85.63%. By harnessing the plastic resistance of steel, the bending resistance of wood can be increased by 32.94%.

Selection of Media for Hardwood Cuttings Container Seedling-raising of Triploid Clones of Populus tomentosa

[Objective] The experiment was aimed to select effective and economical media for container seedling of triploid clones of Populus tomentosa that was carried out. [Method] The sandy loam, peat, perlite, vermiculite, riversand, sludge were taken as media of hardwood cutting and survival rate, seedling height were taken as indexes to select media for container seedling of triploid clones of Populus tomentosa. [Result] Different mixedmedia had great influence on survival rates of container seedlings. Taking peat and vermiculite with the proportion of 5∶2 (M10) or peat ,vermiculite with the proportion of 7∶2 (M11) or sandy loam (M1) as media would generate higher cutting survival rate that was higher than 90.0%. There were significant differece in height increments of container seedlings. Taking sandy loam, peat and vermiculite with the proportion of 6∶2∶2(M5)or sandy loam (M1), seedling height of 60-days the seedling was over 37.0 cm. [Conclusion] According to cost analysis of nursery medium, the optimum medium for hardwood cuttings container seedling-raising of triploid clones of Populus tomentosa was sandy loam.

Study on Hardwood Cutting Propagation Technology of Fraxinus

[Objective] Hard-branch cutting propagation technology of Fraxinus was studied under different environmental conditions. [Method] An orthogonal test was designed based on 1-year-old clones of F. pennsylvanica Marsh Lula 5. Under dif- ferent growth hormones, hormone concentrations and processing time, F. velutina Lula 2 and F. pennsy/vanica Marsh Yuanla 2 were treated by vacuum pump for 10 min, [Result] The decreasing order of cutting survival rates of Lula 5 under different etnvironments was small plastic flowerpot （94.4%）, nutrition cup （92.8%）, sand tray （56.1%）, and cuffings treated with 100 mg/L ABT, for 2 h, 200 mg/L ABT, for 4 h and 100 mg/L IBA for 6 h under large nutrition cup environment grew the best, showing a survival rate of 100%. Lula 2 and Yuanla 2 were subjected to vacuum pump treatment, the survival rate of cuttings under the sand tray environment was averagely 29%; and under the large plastic flowerpot environment, the survival rate of cuttings was averagaly 64.4%, and cuttings of Lula 2 treated with vacuum pump for 10 min in 200 mg/L ABTI showed a survival rate up to 95%, which was the highest among all treatments. It could be seen that whether cuttings were treated with vacuum pump, the survival rate of cuttings in sand tray was lower than those under other environments, while there was no big difference in survival rate of cut- tings between the nutrition cup and plastic flowerpot environments. [Conclusion] This study is of great significance to improvement of cutting propagation coefficient.

Intra-and inter-species variations in carbon content of 14 major tree species in Northeast China

Reducing greenhouse gas emissions is one of the major challenges in combating global warming.Carbon,including in the form of carbon dioxide(CO_(2)),is considered an essential greenhouse gas under human control to demonstrate success in emission reductions.However,many carbon stock quantifications in forest ecosystems still rely on the estimated 50%carbon content instead of more precise species-,tissue-and site-specific values.Thus,this study aimed to thoroughly measure and analyze the carbon content and variability using the 14 major tree species in Northeast China.Over 600 trees were destructively sampled from three different major mountainous regions(i.e.,the Changbai,Daxing’an,and Xiaoxing’an mountains),and the carbon contents of each species were precisely measured to the sub-tissue level.Carbon contents varied significantly between species,with foliage carbon mostly found to be the highest,while root carbon contents were the lowest.Average carbon contents can be ranked as:Ulmus laciniata(43.4%)<Phellodendron amurense(43.5%)<Acer mono(43.8%)<Tilia amurensis(44.2%)<Populus davidiana(44.5%)<Fraxinus mandshurica(44.7%)<Juglans mandshurica(44.9%)<Quercus mongolica(45.3%)<Betulla davurica(45.8%)<Betulla platyphylla(46.7%)<Picea koreansis(46.9%)<Larix gmelinii(47.4%)<Pinus koreansis(48.3%)<Abies nephrolepis(48.3%).Carbon contents were higher in conifers(47.7%)compared to broadleaf species(44.9%).In addition,both tree tissues and growing sites also had a significant effect on carbon content.At the sub-tissue level,only stem’s sub-tissues(i.e.,bark,heartwood,and sapwood)carbon contents showed significant variations.The results suggest that bark should be separated from other stem sub-tissues and considered separately when determining carbon stocks.This research contributes to improving estimates of terrestrial carbon quantifications,and in particular,the values obtained can be used in China’s National Forest Inventory.

Anatomical Properties of Three Lesser Utilised Ghanaian Hardwood Species

Wood is composed of mostly hollow, elongated, spindle-shaped cells that are arranged parallel to each other along the trunk of a tree. The characteristics of these fibrous cells and their arrangement affect strength properties, appearance, resistance to penetration by water and chemical solutions, resistance to decay and many other properties. The characterisation of wood helps in identifying them. In this work, we studied the anatomical properties of three lesser utilised Ghanaian hardwood species namely Albizia ferruginea (Guill. & Perr.) Benth, Blighia sapida K. D. Koenig and Sterculia rhinopetala K. Schum using the light microscope and scanning electron microscope (SEM). Anatomical features studied were fiber length, double fiber wall thickness, fiber proportion, vessel diameter and proportion, rays and axial parenchyma proportions. We observed that the use of SEM in studying the anatomical or ultra-structural aspects of wood gave a clearer understanding of the features and structures found in wood. Anatomical features such as presence of crystals and absence of axial parenchyma in Blighia sapida and the thick wall fibers of Sterculia rhinopetala were better understood.

Forest composition and red oak (Quercus sp.) response to elevation gradients across greentree reservoirs

Elevation gradients within forested wetlands have long been recognized for their role in defining species composition through factors such as hydrology and soil characteristics.Greentree reservoirs(GTRs)are leveeimpounded tracts of bottomland hardwood forest flooded throughout the winter months to provide habitat for overwintering waterfowl.Artificial flooding of GTRs alters the forest composition due to flood frequency,depth,and duration in combination with slight changes in topography.To evaluate the effect of elevation gradients,soil properties,and management techniques in the overstory species composition and red oak(Quercus spp.)species abundance,we inventoried 662 plots across 12 independent GTRs in eastern Arkansas.In the lower elevations ranging from 50.98 to 54.99 m above sea level,the importance value index(IVI)was highest for nuttall oak(Quercus texana)and overcup oak(Quercus lyrata),whereas IVI shifted to cherrybark oak(Quercus pagoda)in the higher elevations ranging from 54.99 to 58.00 m.Alpha diversity did not differ by elevation gradient,soil property,or management technique within GTRs.Beta diversity,using non-metric multi-dimensional scaling(NMDS)analysis,indicated site-specific variability significantly correlated with the environmental predictors,including elevation(R^(2)=0.57),easting(R^(2)=0.47),soil texture(R^(2)=0.21),and pH(R^(2)=0.12).Red oak species-specific mixed-effects modeling of abundance response using Poisson distribution suggested an inverse correlation of nuttall oak and a direct correlation of cherrybark oak abundance with elevation.However,willow oak(Quercus phellos)abundance was not significantly affected by elevation but was by silt loam soil texture and restoration management techniques.These findings will aid management efforts to reduce the dominance of less desirable species that are prominent under specific environmental conditions and promote the dominance of more desirable species.Ultimately GTR sustainability is increasingly important amid the unpredictable impacts of climate change on the preferred red oak species that are economically,ecologically,and environmentally valuable to the sustaining economy of the local community and managing habitats for wildlife.

Influence of individual tree characteristics,spatial structure and logging history on tree-related microhabitat occurrence in North American hardwood forests

Background:Tree-related microhabitats(hereafter,"TreMs")are key components of forest biodiversity but they are still poorly known in North American hardwood forests.The spatial patterns of living trees bearing TreMs(hereafter,"TreM-trees")also remain to be determined.As logging practices can lead to a loss of TreM-trees and of their associated biodiversity,it is essential to identify the factors explaining TreM occurrence to better integrate them into forest management.We therefore inventoried TreMs in 40.5-ha survey strips in northern hardwood forests in Quebec,Canada,while recording the spatial location of each tree.Two strips were located in unmanaged oldgrowth forests,and 2 were in forests managed under selection cutting.All 4 stands were dominated by sugar maple(Acer saccharum Marsh.)and American beech(Fagus grandifolia Ehrn.).Beech bark disease,an exotic pathology,was observed in all the strips.Results:Large diameter at breast height and low tree vigor were the main characteristics explaining the presence of TreMs at the tree scale.TreM-trees presented slight spatial aggregation patterns.These aggregates,however,were not well-defined and were generally constituted by a large number of trees bearing few different types of TreMs.Two TreM classes(broken branch or top and woodpecker lodge)also presented a spatial aggregation.Logging practices had no significant effect on TreM occurrence.Beech bark disease increased the frequency of senescent beeches.The impact of this pathology on TreMs was however mitigated by the small size of infected trees and probably by the short time elapsed since its appearance.Conclusion:The factors explaining the presence and abundance of TreMs on trees has so far been little studied in North American hardwood forests.Our results highlight that TreM-tree characteristics in the surveyed forests are consistent with those of previous studies conducted in other forest types and regions(e.g.,Europe or Northwestern America).To our knowledge,this study is also the first to identify a spatial aggregation of TreM-trees and of specific TreM classes.It will be nevertheless necessary to determine whether the small impact of logging activities we observed results from current or past management practices.

Eight decades of compositional change in a managed northern hardwood landscape

We analyzed over 8 decades of change in forest composition(represented by species proportion of basal area)and size class from more than 400 permanent plots located on the Bartlett Experimental Forest in the White Mountains of New Hampshire.These data represent one of the longest-term landscape-scale records of forest change based on permanent plots in North America.We analyzed the plots based on elevation class,land type indicating assumed successional direction(grouped into coniferous and deciduous),and inventory period within managed and unmanaged portions of the forest.An ongoing shift from small-to large-diameter stems is clear across all species,in response to the overall aging of the forest following exploitative harvesting in the 19th century.Major compositional changes include a continuing decline in shade-intolerant species(paper birch and aspen),along with the mid-tolerant yellow birch.An increase in red maple abundance through the early 1990s has leveled off or reversed.Among shade-tolerant species,increases in beech and red spruce were largely consistent with assumed land type on unmanaged plots,but heavy marking against diseased beech on managed plots restricted increase of that species.Sugar maple declined in abundance except where silvicultural intervention helped maintain it.By contrast,eastern hemlock showed a continuing expansion at all elevations below 600 m.The data continue to show little or no evidence of upward migration of species,despite evidence of recent regional change in climate.However,the BEF is poised for substantial changes when emerald ash borer and hemlock woolly adelgid,both of which are known to infest nearby areas,do arrive.

Stand age structural dynamics of conifer, mixedwood, and hardwood stands in the boreal forest of central Canada

To study the effects of stand development and overstory composition on stand age structure, we sampled 32 stands representing conifer, mixedwood, and hardwood stand types, ranging in ages from 72 to 201 years on upland mesic sites in northwestern Ontario. We defined the stages of stand development as: stem exclusion/canopy transition, canopy transition, canopy transition/gap dynamics, and gap dynamics. Stand age structure of conifer stands changed from bimodal, bimodal, reverse-J, and bimodal, respectively, through the stages of stand development. Mixedwood and hardwood stands revealed similar trends, with the exception of missing the canopy transition/gap dynamic stage in mixedwoods. Canopy transition/gap dynamic stage in hardwoods showed a weaker reverse-J distribution than their conifer counterparts. The results suggest that forest management activities such as partial and selection harvesting and seed-tree systems may diversify standard landscape-level age structures and benefit wildlife, hasten the onset of old-growth, and create desired stand age structures. We also recommend that the determination of old-growth using the following criteria in the boreal forest: 1) canopy breakdown of pioneering cohort is complete and stand is dominated by later successional tree species, and 2) stand age structure is bimodal, with dominating canopy trees that fall within a relatively narrow range of age and height classes and a significant amount of understory regeneration.

Resilience of Bottomland Hardwood Stands Following Agricultural Use on the Santee Experimental Forest

Resilience is a key function that affects an ecosystem’s ability to recover from disturbance. Understanding the extent to which forest communities recover after a long period of disturbance without direct intervention is important to provide context for considering ecosystem response to disturbance regimes. Species composition and structure were recorded on bottomland hardwood stands that were once inland and freshwater tidal rice fields. We sampled 17 old-field sites and 7 reference sites across three geomorphic settings. The old-field sites ranged from 30 to 120 years since agricultural abandonment. A total of 89 species were found across the old field sites and reference sample areas. Of that total, trees comprised 33 species, shrubs—5 species, and vine/herbs/forbs—51 species. Using field data, combined with stand inventory records, aerial photography (1936-2010), and high-resolution LiDAR imagery, we chronicled the evolution of the forest since the cessation of agriculture. Our findings demonstrate how Pinus taeda seeded directly after the rice fields went to fallow;and this conversion of bottomland swamp to rice to pine was a direct result of water management embankments constructed across the landscape to aid in crop irrigation. The remnant water management features may still alter flooding patterns thereby affecting development of Taxodium distichum and Nyssa aquatica in the old-fields. These results suggest, that over the 100+ years forest stands on the Santee Experimental Forest have developed to represent bottomland hardwoods characteristic of the southeastern United States coastal plain.

Rehabilitating Degraded Hardwood Stands on a Bottomland Terrace Site with Overstory Removal and Oak Enrichment Planting: 14-Year Results

Exploitative harvesting can lower stand quality in the short term and diminish forest productivity over the long term. In 2003, a rehabilitation experiment was installed in a southern hardwood stand on a bottomland terrace site, degraded by periodic exploitative cutting, to test the effectiveness of overstory removal and oak enrichment planting for improving stand quality and composition. Overstory removal treatments included clearcutting, stand improvement partial cutting, and an uncut control. Overstory treatment units were either planted with Nuttall oak (Quercus texana) seedlings or not planted. We revisited this study in 2017 to assess the outcome of oak enrichment planting across levels of overstory removal. Results in year 14 indicated poor survival of planted oaks under all overstory removal treatments (14% - 24%) and minimal height growth (0.3 m in 13 years) in areas treated with partial cutting and in uncut areas. Growth performance of planted oaks was significantly enhanced by clearcutting (p < 0.01). However, the overwhelming response of natural oak regeneration initiated by clearcutting trumped the contribution from enrichment planting on this low quality bottomland terrace site. Enrichment planting of Nuttall oak did not increase oak regeneration success in our study. The ineffectiveness of enrichment planting in this study was likely related to the shady understory of partially cut stands and intense competition in clearcuts. Additional treatments, such as pre-planting site preparation and post-planting release may be necessary for enhancing and maintaining competitiveness of planted red oak seedlings on similar sites.

Predicting Stem Windthrow Probability in a Northern Hardwood Forest Using a Wind Intensity Bio-Indicator Approach

Unlike fire or insect outbreaks, for which a suppression program can be implemented, it is impossible to prevent a windstorm event or stop it while it is occurring. Reducing stand susceptibility to windstorms requires a good understanding of the factors affecting this susceptibility. Distinct species- and size-related differences in stem windthrow susceptibility are difficult to obtain because it is impossible to distinguish their relative effects from those of wind intensity. Using a damage assessment database (60 20-metre radius plots) acquired after an exceptional wind storm in Western Quebec in 2007, we developed an approach in which proportions of windthrown sugar maple poles were used as bio-indicators of wind intensities affecting the plots. We distinguished between single and interactive effects of wind intensity, species, stem size, and local basal area on stem windthrow susceptibility. The best logistic regression model predicting stem windthrow included the wind intensity bio-indicator, species, basal area, and the species by diameter at breast height (DBH, 1.3 m) interaction. Stem windthrow probability generally increased with DBH and decreased with basal area. Species wind-firmness was ordered as: yellow birch > sugar maple = eastern hemlock = American beech > ironwood > basswood = other hardwoods = other softwoods. Our method remained an indirect method of measuring wind intensity and its real test would require a comparison with anemometer measurements during a windstorm. Despite its indirect nature, the method is both simple and ecologically sound. Hence, it opens the door to conducting similar windthrow studies in other ecosystems.

Behavior of wood basic density according to environmental variables

The relationships between climate conditions and wood density in tropical forests are still poorly understood.To quantify spatial dependence of wood density in the state of Minas Gerais(MG,Brazil),map spatial distribution of density,and correlate density with climate variables,we extracted data from the Forest Inventory of Minas Gerais for 1988 trees scaled throughout the territory and measured wood density of discs removed from the trees.Environmental variables were extracted from the database of the Ecological-Economic Zoning of Minas Gerais.For spatial analysis,tree densities were measured at 44 georeferenced sampling points.The data were subjected to exploratory analysis,variography,cross-validation,model selection,and ordinary kriging.The relationships between wood density and environmental variables were calculated using dispersion matrices,linear correlation,and regression.Wood density proved to be highly spatially dependent,reaching a correlation of 96%,and was highly continuous over a distance of 228 km.The distribution of wood density followed a continuous gradient of 514-659 kg m^(−3),enabling corre-lation with environment variables.Density was correlated with mean annual precipitation(−0.57),temperature(0.63),and evapotranspiration(0.83).Geostatistical methods proved useful in predicting wood density in native tropical forests with different climate conditions.Our results confirmed the sensitivity of wood density to climate change,which could affect future carbon stock in forests.

Economic analysis on log damage during logging operation in Caspian Forests

Waste wood was studied in an economic enterprise by logging, function, tree species and log size in four Caspian hardwood sites. Dam- aged logs were recorded with additional information obtained for the location, dimensions and type of damage. The data were analyzed statis- tically to determine significant differences of damage during logging process. The results indicated that animal harvesting systems cause more volume （40.5% of log volume） and value loss （89.5 $.m^-3） to logs than mechanized harvesting systems （13.9% and 6.0 $.m^-3）, also bucking resulted in significantly more volume （9.9% of log volume） and value loss （5.5 $.m-3） when compared to skidding （0.2% of log volume and 0.2 $.m^-3）, decking （0.4% of log volume and 0.2 $.m^-3） and loading （0.2% of log＇s volume and 0.3 $.m^-3） operations. Study showed that the processes of skidding, decking and loading of logs have very little impact on dam- age levels. Volume and value losses of damaged logs are not sensitive to tree species and log size. The information from the field study is impor- tant in creating new guidelines or training to help minimize hardwood log damage during the timber harvesting process.

Evidence of Dual Scale Porous Mechanisms During Fluid Migration in Hardwood Species (Ⅱ) A Dual Scale Computational Model to Describe the Experimental Results

The second part of this paper is devoted to the computational modelling of transient water migration in hardwood. During re-saturation, the moisture content, measured during the process by using X-ray attenuation (see part 1 of this paper), increases quickly very close to the cavity, but requires a very long time for the remaining part of the sample to absorb the moisture in wetting. For this configuration and this material, the macroscopic approach fails. Consequently, a dual-porosity approach is proposed. The computational domain uses a 2-D axisymmetric configuration for which the axial coordinate represents the macroscopic longitudinal direction of the sample whereas the radial coordinate allows the slow migration from each active vessel towards the fibre zone to be considered. The latter is a microscopic space variable. The moisture content field evolution depicts clearly the dual scale mechanisms:a very fast longitudinal migration in the vessel followed by a slow migration from the vessel towards the fibre zone.The macroscopic moisture content field resulting from this dual scale mechanism is in quite good agreement with the experimental data.

SIMULATION OF SOIL NITROGEN MINERALIZATION AND NITRIFICATION IN TWO NORTHERN HARDWOOD FOREST ECOSYSTEMS

A process-based, biological model is presented that simulates soil nitrogen (N) mineralization and nitrification in two northern hardwood forest ecosystems in the Upper Peninsula of Michigan. The soil system is divided into two compartments (forest floor and mineral soil) since quantity and quality of the organic substrate, and the important driving variables (temperature and moisture) for the model vary between these two compartments. The model focuses on the central position of microorganisms in the N mineralization and nitrification processes, and the use of multiplicative factors to account for the effect of temperature, moisture and carbon(C):N ratio on these processes.The model has been validated with data from two northern hardwood stands in the Upper Peninsula of Michigan. A close agreement between calculated and observed monthly means was obtained in both stands, especially for net N mineralization, which plays a very impormnt role in determining available N. The nitrification rates had relatively larger variation than the N mineralization rates, but the model adequately described the seasonal trends of the observed values. A simple sensitivity analysiwas performed to assess the response of the model to changes in important variables (temperature, moisture, organic N, and C:N ratio) between the two study sites. This analysis showed that increased temperature and higher organic N levels consistently increased N mineralization and nitrification in the both stands. The model's results were most sensitive to moisture changes in forest floor, but were not sensitive to moisture changes in the mineral soil. In contrast, C:N ratio was influential in the mineral soil, but did not have any effect in the forest floor.

Mechanical Characterisation of Densified Hardwood with Regard to Structural Applications

The demand for high-performance,yet eco-friendly materials is increasing on all scales from small applications in the car industry,instrument or furniture manufacturing to greater dimensions like floorings,balcony furnishings and even construction.Wood offers a good choice on all of these scales and can be modified and improved in many different ways.In this study,two common European hardwood species,Beech(Fagus sylvatica L.)and Ash(Fraxinus excelsior L.)were densified in radial direction by thermo-mechanical treatment and the densified product was investigated in an extensive characterisation series to determine all relevant mechanical properties.Compression in the three main directions(longitudinal,tangential,radial)and tension perpendicular to the grain(tangential,radial)were tested and compared to reference specimens with native density.Strength and modulus of elasticity were determined in all tests.In addition,a Life Cycle Assessment was carried out to evaluate the environmental impact associated to the densification process.The experimental investigations showed that strength and stiffness of hardwood in the longitudinal and tangential directions improve significantly by radial densification,whereas some properties in the radial direction decrease.The Life Cycle Assessment showed that artificial wood drying has higher impact than wood densification.Furthermore,the transport distance of the raw material highly influences the environmental impact of the final densified product.The paper then also offers an overview of possible applications in structural timber construction.Densified hardwood is a viable option as local reinforcement,where high compressive or tensile strength is needed.The wood densification process offers an alternative to the use of carbon-intense steel components or hardwoods from tropical forests.

Verification of Depth Adjustment Factor in Eurocode 5 for Tropical Hardwood Timbers

The depth adjustment factor for bending strength stated in Eurocode 5(EC5)is only applicable to timbers having a characteristic density below 700 kg/m^(3).However,most Malaysian timbers are hardwood,some with a characteristic density reaching above 700 kg/m^(3).Therefore,the objective of this study was to examine whether the depth adjustment factor stipulated in EC5 is valid for Malaysian hardwood timbers.Six timber species were selected for this study,namely Kapur(Dryobalanops C.F.Gaertn.),Kempas(Koompassia Maingay ex Benth.),Keruing(Dipterocarpus C.F.Gaertn.),Light red meranti(Shorea Roxb.ex C.F.Gaertn.),Geronggang(Cratoxylum Blume)and Balau(Shorea Roxb.ex C.F.Gaertn.).The determination of bending strength and characteristic density was conducted according to BS EN 408:2010 and BS EN 384:2016,respectively.A graph for mean bending strength vs.(150/h)was plotted for each timber species.The power function was selected to analyze the relationship between the two variables.The power of the regression equations varied depending on the characteristic density of the timber species.For species with a characteristic density below 700 kg/m^(3),such as Kapur,Keruing,and Light red meranti,the power was between 0.16 to 0.17.In contrast,for species having a characteristic density above 700 kg/m^(3),namely Kempas and Balau,the power was higher at 0.23 and 0.24,respectively.Geronggang was an exception to this pattern.These values are close to the depth adjustment factor given in EC5,which is 0.2.Based on the results,it can be suggested that the adjustment factor of 0.2 is also applicable to Malaysian hardwood timbers with a characteristic density above 700 kg/m^(3).

Herbaceous-Layer Community Dynamics along a Harvest-Intensity Gradient after 50 Years of Consistent Management

In 1958, a demonstrational cutting trial totaling 22.2 ha was established in a northern hardwood forest in Alberta, MI. Eight different treatments were installed, including four diameter-limit treatments (56 cm, 41 cm, 30 cm, and 13 cm), three single-tree selection treatments with residual basal areas of 21 m2·ha–1, 16 m2·ha–1, and 11 m2·ha–1, and an uncut control. Within each treatment, a 0.4-ha permanent plot was established and subdivided into 0.04-ha square subplots. Harvests have been implemented every ten years with the most recent harvest occurring during the winter of 2008 - 2009. We quantified ground layer vegetation response before and after the most recent harvest. Nonmetric multidimensional scaling (NMS) ordination showed a very distinct separation between the most intensive management treatment (13-cm diameter-limit treatment) and the uncut control. Compositionally, the diameter-limit treatments moved with greater directionality and magnitude towards the 13-cm diameter-limit treatment following harvest, while compositional change in the residual basal area treatments was less pronounced and lacked strong directionality. Herbaceous species percent cover generally decreased with increasing residual overstory basal area across treatments. Weedy and early successional species were most abundant under lower residual basal area and diameter-limit treatments. Results based on 50 years of continuous management suggest that diameter-limit harvests likely have a greater impact on the herbaceous community than single-tree selection or no management.