Early-diverging Titanosauriform(Dinosauria,Sauropoda)Teeth from the Lower Cretaceous Yixian Formation of Southeastern Inner Mongolia,Northeast China

Three eusauropod teeth(SDUST-V1064,PMOL-AD00176,PMOL-ADt0005)are reported from the Lower Cretaceous Yixian Formation of Ningcheng,southeastern Inner Mongolia,China.Two of them(SDUST-V1064,PMOL-AD00176)are assigned to early-diverging titanosauriforms in having slightly mesiodistal expansion at the base of the tooth crown,a slenderness index value>2.0 and<4.0,and D-shaped cross section.Furthermore,SDUST-V1064 and PMOL-AD00176 are referred as an Euhelopus-like titanosauriform on the basis of having a sub-circular boss on the lingual surface and an asymmetrical crown-root margin which slants apically,respectively.CT scan data of SDUST-V1064 reveals new dental information of early-diverging titanosauriforms,for example,the enamel on the labial side thicker than that on the lingual side,an enamel/dentine ratio of 0.26 and a boss present on the lingual side of the dentine of the crown.

Hydrocarbon accumulation history in Lower Cretaceous in northern slope of Bongor Basin in Chad,Central Africa

Based on the analysis of the fluid inclusion homogenization temperature and apatite fission track on the northern slope zone of the Bongor Basin in Chad,this paper studied the time and stages of hydrocarbon accumulation in the study area.The results show that:(1)The brine inclusions of the samples from the Kubla and Prosopis formations in the Lower Cretaceous coexisting with the hydrocarbon generally present two sets of peak ranges of homogenization temperature,with the peak ranges of low temperature and high temperature being 75–105℃ and 115–135℃,respectively;(2)The samples from the Kubla and Prosopis formations have experienced five tectonic evolution stages,i.e.,rapid subsidence in the Early Cretaceous,tectonic inversion in the Late Cretaceous,small subsidence in the Paleogene,uplift at the turn of the Paleogene and Neogene,and subsidence since the Miocene,in which the denudation thickness of the Late Cretaceous and after the turn of the Paleogene and Neogene are~1.8 km and~0.5 km,respectively.The cumulative denudation thickness of the two periods is about 2.3 km;(3)Using the brine inclusion homogenization temperature coexisting with the hydrocarbon as the capture temperature of the hydrocarbon,and combining with the apatite fission track thermal history modeling,the result shows that the Kubla and Prosopis formations in the Lower Cretaceous on the northern slope of the Bongor Basin have the same hydrocarbon accumulation time and stages,both of which have undergone two stages of hydrocarbon charging at 80–95 Ma and 65–80 Ma.The first stage of charging corresponds to the initial migration of hydrocarbon at the end of the Early Cretaceous rapid sedimentation,while the second stage of charging is in the stage of intense tectonic inversion in the Late Cretaceous.

Dongbeititan dongi,the First Sauropod Dinosaur from the Lower Cretaceous Jehol Group of Western Liaoning Province,China

We herein describe a partial postcranial skeleton of a sauropod dinosaur recovered from the Lower Cretaceous Jehol Group in the Beipiao area of western Liaoning Province, northeastern China. A suite of features it possesses, including the camellate internal structure of its presacral elements, the existence of pneumatocoels on the proximal ends of the dorsal ribs, and especially the medially deflected proximal portion of the femur, definitively establish the titanosauriform affinities of the specimen. It differs from other titanosauriforms in having a craniocaudally elongate coracoid with a squared cranioventral extreme and a long, smooth, and slightly convex acetabular edge of the pubis. It represents a new taxon, Dongbeititan dongi gen. et sp. nov. Comparative studies suggest that Dongbeititan is a basal titanosauriform, more derived than Euhelopus, Fusuisaurus, and Huanghetitan, but less derived than Gobititan and Jiutaisaurus. Dongbeititan represents the first sauropod dinosaur reported from the Lower Cretaceous Jehol Group of western Liaoning Province.

A New Species of Cathayornis from the Lower Cretaceous of Inner Mongolia,China and Its Stratigraphic Significance

An incomplete postcranial avian skeleton is described from the Lower Cretaceous Jingchuan Formation of Otog Banner of western Inner Mongolia and referred to a new species of Cathayornis, C. chabuensis sp. nov. This is the first report of a Cathayornis from outside Liaoning Province. The new discovery indicates that Cathayornis coexisted with Otogornis genghisi, and a more detailed comparison between these two enantiornithine genera shows that Otogornis represents a more primitive genus than Cathayornis. Our analyses further indicate that Cathayornis is an arboreal bird. The discovery of a Cathayornis from this region also confirms that the avian fossil-bearing Jingchuan Formation is comparable to the Jiufotang Formation of the upper Jehol Group in western Liaoning, and should be referred to the middle-late Early Cretaceous.

Precious Fossil-Bearing Beds of the Lower Cretaceous Jiufotang Formation in Western Liaoning Province, China

Based on the subdivision into three members of the Lower Cretaceous Jiufotang Formation in western Liaoning, this paper deals mainly with the division and correlation of precious fossil birdand reptile-bearing beds of the formation in the Dachengzi, Chaoyang, Dapingfang.Meileyingzi and Fuxin-Yixian basins. Among them, the precious fossil-bearing beds in the Dachengzi Basin may be recognized as the Xidagou Bed of the second member and the Yangcaogoudonggou Bed of the third member; those in the Chaoyang Basin may be confirmed as the Shangheshou Bed of the second member and the Dongpochi Bed of the third member; those in the Dapingfang-Meileyingzi Basin are as the Lamagou Bed of the second member, the Huanghuagou Bed of the lower third member and the Yuanjiawa Bed of the upper third member; and those in the Fuxin-Yixian Basin are listed as the Tuanshanzi Bed of the second member and the Pijiagou Bed of the third member. Since these basins are distinctly separated and the bird and reptile fossils are mostly new genera and species, we have to use the associated fossil ostracod assemblages as index to correlate the Xidagou Bed with the Shangheshou Bed and the Lamagou Bed, and to correlate the Yuanjiawa Bed with the Yangcaogoudonggou Bed and the Pijiagou Bed. Primarily, we established the sequence of the precious fossil-bearing beds of the Jiufotang Formation in western Liaoning. They are represented by, in ascending order, the Xidagou Bed of the second member, the Huanghuagou Bed of the lower third member, and the Yuanjiawa Bed of the upper third member. Obviously, this work has significance for the study on the time-space distribution and radiation of birds and dinosaurs during the period of the middle Early Cretaceous.

An Adult Specimen of Hongshanosaurus houi (Dinosauria:Psittacosauridae) from the Lower Cretaceous of Western Liaoning Province, China

A new specimen consisting of a complete skull and lower jaw was discovered from the Lujiatun bed of the Lower Cretaceous Yixian Formation in the Lujiatun site, Beipiao, western Liaoning Province. It represents an adult specimen of Hongshanosaurus houi. Comparative studies show that Hongshanosaurus is distinguishable from its sister taxon Psittacosaurus by having the preorbital portion about half of the basal skull length, and the elliptical external naris, orbit, and a lower temporal fenestra oriented caudodorsally.

A New Species of Huaxiapterus (Pterosauria: Pterodactyloidea) from the Lower Cretaceous of Western Liaoning, China with Comments on the Systematics of Tapejarid Pterosaurs

A new species of tapejarid pterosaur, Huaxiapterus coroUatus sp. nov. is erected on the basis of a nearly complete skull and postcranial skeleton from the Lower Cretaceous Jiufotang Formation of Liaoning Province, China. Huaxiapterus coroUatus sp. nov. is characterized by a hatchet-shaped rectangular process on the premaxilla, whose short axis is perpendicular to the anterior margin of the premaxillae. Except for this process, other characters of the skull such as the breadth of the snout between the anterior margin of the nasoantorbital fenestra and the anterior margin of the premaxilla are similar to that of Huaxiapterus jii. Huaxiapterus and a second Chinese tapejarid, Sinopterus, share several unique cranial characters in common with Tapejara and these three genera appear to be more closely related to each other than to other azhdarchoids. The Chinese tapejarids （Sinopterus and Huaxiapterus） have relatively elongate skulls and weakly developed cranial crests and seem to be less derived than Tapejara, with its shorter, deeper skull and large cranial crest. Tupuxuarids （Tupuxuara and Thalassodromeus） have often been associated with tapejarids in the family Tapejaridae, but this relationship is controversial because some phylogenetic analyses have supported the pairing of tupuxuarids with Azhdarchidae. We propose that Tapejaridae be restricted to Tapejara, Sinopterus and Huaxiapterus.

On the Lower Cretaceous in Yixian County of Jinzhou City,Western Liaoning,China

Traditionally, the lacustrine deposits in Yixian County of Western Liaoning have been assigned to either ＂the Dakangpu Bed＂ of the Yixian Formation or the Jiufotang Formation. However,a 206pb/238U age of 126.47±0.87 Ma is newly obtained from the tuff sample （06051） of the lacustrine deposits in the Baitaigou fossil site of Potaizi village, Yixian County, indicating that the lacustrine deposits in Potaizi area are approximately equivalent to the Wumingshan volcanics of the Sihetun Bed of the Yixian Formation in Beipiao. From our new dating data have arisen some new problems： Where could we find the Jiufotang Formation in Yixian County？ How widely distributed is the Jiufotang Formation in Western Liaoning？ Could the traditionally named Jiufotang Formation somewhere in Western Liaoning be the synchronous deposits of other litho-units （e.g. the Yixian Formation） in different facies？ Could some lacustrine deposits somewhere in Western Liaoning have been assigned to the Jiufotang Formation？

Sauropod Teeth from the Lower Cretaceous Luohandong Formation of Ordos Basin,Inner Mongolia

The Early Cretaceous Zhidan Group in the northern Ordos Basin, Inner Mongolia, yielded a large number of tetrapods, including turtles, choristoderes, crocodyliforms, psittacosaurs, stegosaurs, theropods and birds. Well-preserved sauropod teeth have been found in the Luohandong Formation, a middle-upper unit of the Zhidan Group. The large V-shaped wear facet, low slenderness index value, labial grooves, lingual ridge and concavity on the tooth crown suggest that these teeth are from titanosauriforms. Moreover, the presence of the prominent bosses on the lingual side of the tooth crown indicates these teeth should be identified as Euhelopus teeth further. The existence of Euhelopus in Ordos Basin （Inner Mongolia）, Shandong Province and western Liaoning Province shows some connections about vertebrate faunas during Early Cretaceous in these areas. Other tetrapod groups such as turtles （Sinemys, Ordosemys）, choristoderes （Ikechosaurus）, psittacosaurs （Psittacosaurus） and birds （Cathayornis） provide more evidences for this viewpoint.

Palynological Analysis of Amber-Bearing Clay from the Lower Cretaceous of Central Lebanon

An amber-bearing lignitic layer of sandy clay from the Lower Cretaceous of Central Lebanon （Mderej-Hammana）yielded a well-preserved,moderately variegated palynoflora,which origin is mixed between land plants and marine microflora.Its detailed analysis led to fulfill its inventory,to propose a paleoenvironmental reconstruction,and to draw the paleoclimate which prevailed over the region：an estuarian area under a rather humid,temperate climate;a variety of ferns grew near the shore-side and in the inward land.A tiny piece of amber containing angiospermous pollen grains of stratigraphical interest allows a precise dating.The marine microflora,poorly diversified,includes chitinous foraminifer linings and dinoflagellate cysts,among which Early Aptian guide taxa are present;their occurrence slightly narrows the stratigraphical range indicated by some palynological taxa which are related to land plants.

A New Understanding of the Sedimentary Environment of the Laiyang Group in the Lower Cretaceous of Lingshan Island,Shandong Province,East China

The sedimentary background and sedimentary environment of the Lower Cretaceous of Lingshan Island,Shandong,remain unclear.There is considerable controversy as to whether the sedimentary environment was marine or continental.In this study,analyses were conducted with respect to indicators of paleosalinity,the chemical composition of clay,paleobiota,the characteristics of strata exhibiting salinization in eastern China during the Early Cretaceous,and the relationship between paleosalinity and paleoclimate.The results indicate that the sedimentary environment of the Lower Cretaceous Laiyang Group of Lingshan Island was that of a salified lake rather than a marine environment.This study not only provides a reference for determining the sedimentary environment of the Lower Cretaceous Laiyang Group of Lingshan Island but also offers a new perspective for the study of Cretaceous strata with salinization characteristics in eastern China.

Large-Sized Ankylosaur(Dinosauria) from the Lower Cretaceous Jiufotang Formation of Western Liaoning,China

Two incomplete bones referred to ankylosaur scapulocoracoid and humerus are described, which were discovered from the Early Cretaceous Jiufotang Formation of Dachengzi Town, Kazuo County, Liaoning Province. Ankylosaur dinosaurs have ever been found in the Lower Cretaceous Yixian, Fuxin and Sunjiawan Formations of western Liaoning. The occurrence of ankylosaurs within Jiufotang Formation improves our knowledge on the ankylosaur stratigraphic distribution. Based on the length of scapulocoracoid, the new dinosaur is estimated to exceed 6.0 meters long, representing the largest ankylosaur in western Liaoning.

A New Species of Cupressinocladus from the Lower Cretaceous of Guyang Basin,Inner Mongolia,China and Cluster Analysis

Cupressinocladus Seward is a fossil genus of conifers and conifer fossils with reproductive organs are very rare. In general, it is difficult to understand the natural affinities with other conifers. In this paper, a new species, Cupressinocladus guyangensis P.H. Jin et B.N. Sun sp. nov., is reported based on branches with immature female cones from the Lower Cretaceous Guyang Formation of the Guyang Basin in Inner Mongolia, northern China. The foliage shoots are decussate. Leaves are decussate, imbricate, scale-like, weakly dimorphic, and bear longitudinal glands on the abaxial view. Stomata complexes are haplocheilic, monocyclic, irregularly arranged, and spread along the leaf margin. Immature female cones are subglobose with 6-8 cone scales, and three subglobose ovules arranged in a row at the base of the cone scales. Moreover, we performed cluster analysis using a statistics and machine learning toolbox for 23 fossils and extant species based on 16 morphological characters. The result implies that the new species bears a close resemblance to the extant Cupressusfunebris Endl. and might have nearest systematic affinities to it.

Discovery of Oil and Gas Shows in the Lower Cretaceous of the Hongmiaozi Basin, Southeastern Songlibao Peripheral Basin

Objective The Hongmiaozi Basin in the southeastern part of the Songliao peripheral basin is a new oil and gas exploration area with a very low exploration level.This basin covers an area of 795 km;.It is adjacent to the Liuhe Basin and Tonghua Basin in the north,and to the Huanren Basin in the south.In order to identify the basic oil and gas geological characteristics,the deep structures and the

Talaya batraba gen.et sp.nov.- the First Nymph of a Protopsyllidiid(Hemiptera:Sternorrhyncha:Psyllomorpha) from the Lower Cretaceous Amber of Lebanon

A hemipteran nymph of the sternorrhynchan lineage, placed in the family Protopsyllidiidae is the first found in the fossil record, based on an inclusion in amber from the Lower Cretaceous of Hammana / Mdeyrij, Abeih Formation, Central Lebanon. Based on distinctive features such as a median dorsal elevation and the presence of a large, conical, exposed, setiferous anal tube, the fossil is placed in Talaya batraba gen. et sp. nov. and the newly erected taxon is compared to known nymphs of extinct Protopsyllidiidae. The evolutionary traits of the family and its relatives are considered.

Integrated Biostratigraphy, Depositional Setting and Geochemical Analyses for Petroleum Potential Evaluation of the Lower Cretaceous(Barremian – Albian) Strata of the Koppeh-Dagh Basin, Northeastern Iran

The lower Cretaceous rock units of the Koppeh-Dagh Basin of northeastern Iran were investigated here in terms of biostratigraphy,depositional setting and geochemical analyses to find out if they,alike other parts of the world,are rich in petroleum.For this purpose,a stratigraphic framework is established using calcareous nannofossil and palynological elements.A nannoplankton zonation based on which subzones of the zones CC7–CC8 of Sissingh(1977)and their equivalent NC6–NC8 of Roth(1978)was established indicating a Late Barremian–Albian age.Palynological assemblages led us to establish the local palynozone of Odontochitina operculata.A dominantly marginal basin to a transitional zone between shelf and basin under a dysoxic–anoxic condition with low to moderate sedimentation rates coincided with a gradual sea level rise was introduced as the environment of deposition for the strata via interpretation of the palynological parameters and quantitative palynology.The obtained data from Rock-Eval pyrolysis in compilation with palynofacies analysis reveals that the studied succession contains mainly gas-prone type III kerogen.The Spore Coloration Index(SCI)alongside with the Rock-Eval pyrolysis results(low values of HI and TOC)proves that these rock units locally produced natural gas during the time under consideration.

Sedimentary Facies Models on Carbonatite in the Upper Shuaiba Member of Lower Cretaceous in Daleel Field, Oman

The Upper Shuaiba Member （USH） is the main force pay bed in the Daleel field in northern Oman; 5 layers including A, B, C, D, and E were divided in profile, and layer D and layer E are the main beds. With the development of exploration in the Daleel oil field, studying the sedimentary systems about their inner composition and the collocation in dimension, and setting up the sedimentary models in the USH are becoming more and more necessary and important to meet the further exploration requirement. Based on the data of geology, seism, and paleo-biology, according to the analysis method on carbonatite depositional system, the litho-facies assemblage and sedimentary environment in the USH were studied. Intershoal low-lying sub-facies （where the water depth is 10-50 m） and shallow shoal sub-facies （where the water depth is not more than 10 m） were extinguished in the layer D, and storm deposit was found in layer El, in which intershoal low-lying sub-facies also developed. The feature of the sedimentary sub-facies and the sedimentary condition were analyzed, and the sedimentary model was set up in the article： the carbonatite intershoal low-lying developed under the background of open land in shallow sea, where storm events usually occurred in the Lower Cretaceous in the area.

The Discovery of the Bivalve Fossil Arguniella from the Lower Cretaceous Shangkuli Formation in Northern Greater Khingan Mountains, China: Implications for Stratigraphic Correlation

Objective Typical Early Cretaceous non-marine bivalve fossils from Eastern Asia are represented by the Arguniella-Sphaerium assemblage,and they were important members of the Jehol Biota.Historically,the genus Arguniella in western Liaoning had been mistakenly identified as the Jurassic Ferganoconcha,re-identified as Arguniella with。

The Lower Cretaceous Giant Dyke Swarms in the Arctic and Its Significance for linking of Siberia-Iceland Hot-Spot Track

Siberia-Iceland hot-spot track is the one of hot-spot track preserved on continent.Although this hot-spot track has clear plume"heads"-Siberia large igneous province,owing to thick continental lithosphere,its continuing

Influences of burial process on diagenesis and high-quality reservoir development of deep-ultra-deep clastic rocks:A case study of Lower Cretaceous Qingshuihe Formation in southern margin of Junggar Basin,NW China

Taking the Lower Cretaceous Qingshuihe Formation in the southern margin of Junggar Basin as an example,the influences of the burial process in a foreland basin on the diagenesis and the development of high-quality reservoirs of deep and ultra-deep clastic rocks were investigated using thin section,scanning electron microscope,electron probe,stable isotopic composition and fluid inclusion data.The Qingshuihe Formation went through four burial stages of slow shallow burial,tectonic uplift,progressive deep burial and rapid deep burial successively.The stages of slow shallow burial and tectonic uplift not only can alleviate the mechanical compaction of grains,but also can maintain an open diagenetic system in the reservoirs for a long time,which promotes the dissolution of soluble components by meteoric freshwater and inhibits the precipitation of dissolution products in the reservoirs.The late rapid deep burial process contributed to the development of fluid overpressure,which effectively inhibits the destruction of primary pores by compaction and cementation.The fluid overpressure promotes the development of microfractures in the reservoir,which enhances the dissolution effect of organic acids.Based on the quantitative reconstruction of porosity evolution history,it is found that the long-term slow shallow burial and tectonic uplift processes make the greatest contribution to the development of deep-ultra-deep high-quality clastic rock reservoirs,followed by the late rapid deep burial process,and the progressive deep burial process has little contribution.