Statistical Downscaling Retrieval of Land Surface Temperature in an Area with Complex Landforms in the Eastern Qinling Mountains of China Based on Sentinel-2/3 Satellite Data

The study of land surface temperature(LST)is of great significance for ecosystem monitoring and ecological environmental protection in the Qinling Mountains of China.In view of the contradicting spatial and temporal resolutions in extracting LST from satellite remote sensing(RS)data,the areas with complex landforms of the Eastern Qinling Mountains were selected as the research targets to establish the correlation between the normalized difference vegetation index(NDVI)and LST.Detailed information on the surface features and temporal changes in the land surface was provided by Sentinel-2 and Sentinel-3,respectively.Based on the statistically downscaling method,the spatial scale could be decreased from 1000 m to 10 m,and LST with a Sentinel-3 temporal resolution and a 10 m spatial resolution could be retrieved.Comparing the 1 km resolution Sentinel-3 LST with the downscaling results,the 10 m LST downscaling data could accurately reflect the spatial distribution of the thermal characteristics of the original LST image.Moreover,the surface temperature data with a 10 m high spatial resolution had clear texture and obvious geomorphic features that could depict the detailed information of the ground features.The results showed that the average error was 5 K on April 16,2019 and 2.6 K on July 15,2019.The smaller error values indicated the higher vegetation coverage of summer downscaling result with the highest level on July 15.

Winter diet and digestive tract of the Golden Pheasant(Chrysolophus pictus) in the Qinling Mountains,China

The winter diet and morphological structure of the gastrointestinal tract of the Golden Pheasant(Chrysolophus pictus) was investigated in the Qinling Mountains,Shaanxi Province in 2002/2003.Individual food items in crops were identified by species,where possible,using standard taxonomic methods.The Golden Pheasant consumes exclusively vegetarian foods in the winter,of at least 14 plant species,such as crops and other vegetable species.The digestive tract of the Golden Pheasant is composed of an oesophagus,a stomach,a relatively long intestine measuring 3.4 × standard body length,two fully-developed caeca and a relatively short colon,typical for herbivorous birds.Pebbles of different sizes(0.5-3 mm in diameter) were very frequent in the gizzard.The average dry weight of the pebbles was 10.4±2.5 g and was positively correlated with the weight of digesta in the gizzard(Pearson r = 0.747,p < 0.01,n = 37).The mucosa surface pH of the digestive tract of the Golden Pheasant was slightly acidic,but higher in the crop and gizzard.

GIS-Based Assessment of Land Suitability for Optimal Allocation in the Qinling Mountains, China

A GIS-based method was used to assess land suitability in the Qinling Mountains, Shaanxi Province of China through simultaneous consideration of physical features and current land use. Through interpretation of Landsat TM images and extensive field visits the area was modeled into 40 land types in five altitudinal zones (valleys and gullies, hillsides and terraces, foothills, mid-mountain, and sub-alpine mountain). Then, a suitability score was assigned to five physical factors (climate, hydrology, topography, soil, and vegetation). Next, their integrated overall suitability value scores were compared with the observed land cover to determine whether it should be reallocated a new use. Results showed that the five suitability classes of agriculture, forest, grassland, farmland-woodland, and scrub-pasture had altitudinal stratification and a total of 1151 km2 (8.89%) of lands on the northern slopes of the Qinling Mountains had to be reallocated. To achieve this reallocation, 657 km2 of arable land should be reduced, and forest, grassland and scrub-pasture increased by 615 km2, 131 km2 and 405 km2, respectively. Implementation of these recommended land reallocations should help achieve suitable use of land resources and prevent land degradation.

Determination of the Neoproterozoic Shicaogou Syn-collisional Granite in the Eastern Qinling Mountains and Its Geological Implications

The Shicaogou granite has been identified as a magnesian (Fe-number=0.71-0.76), calcic to calc-alkalic (MALI=3.84-5.76) and peraluminous (ASI=1.06-1.13) granite of the syn-collisional S-type, with high SiO2(>71%), A12O3 (>13%) and Na2O+K2O (6.28%-7.33%, equal for NaO2 and K2O). Trace element and REE analyses show that the granite is rich in LILE such as of Rb, Sr, Ba and Th, and poor in HFSE like Yb, Y, Zr and Hf. Its Rb/Sr ratio is greater than 1; the contents of Nb and Ta, and the ratio of Nb/Ta as well as the REE geochemical features (e.g. REE abundance, visible fractionation of LREE and HREE and medium to pronounced negative Eu anomalies) are all similar to those of crust-origin, continent-continent syn-collisional granite. Moreover, the granite exhibits almost the same pattern as that of the typical continent-continent syn-collisional granite on the spider diagram and all samples fall within the syn-collisional granite field.The cathodoluminescence (CL) investigations have revealed that the zircon from the Shicaogou granite represents a typical magmatic product characterized by its colorless, transparent and euhedral crystals, and distinct zoning of oscillatory bands. Residual cores of irregular zircon can be found in a few enhedral grains. Trace element studies of the zircon grains, with high contents of P, Y, Hf, Th, U and REE and high ratios of Th/U, obviously positive Ce anomalies and HREE enrichment compared to LREE, also result in the same conclusion.The LA-ICP-MS U-Pb isotopic data from 24 spots of 21 zircon grains demonstrate that 20 spots in the oscillatory zone yield an average weighted 206Pb/238U age of 925±11 Ma, indicating that the Shicaogou granite was formed in the Neoproterozoic. Combined with other Neoproterozoic syn-collisional granites found in the study area, the present geochronological determination can further reveal that collision-amalgamation events could have occurred among some continental blocks in the Qinling orogenic belt during the Neoproterozoic. This in turn provides an accurate chronological constraint on the Neoproterozoic break-up and convergence in the belt.

^(40)Ar/^(39)Ar Thermochronological Constraints on the Timing of Collisional Orogeny in the Mian-Lite Collision Belt,Southern Qinling Mountains

Silurian, Devonian and Carboniferous geological bodies in the Mianxian-Lueyang (Mian-Lue) collisional belt (MLB) and its neighbouring areas, southern Qinling Mountains, China, show similar characteristics of having undergone deformation of two stages. The earlier one, which is inferred to be related to collisional orogeny between the Yangtze and Sino-Korean palaeocontinents based on previous geological data, is responsible for large-scale, north-verging recumbent folds and overthrusts, and associated with low greenschist fades metamorphism. 40Ar/39Ar dating of three muscovite samples taken from different localities yields plateau ages of 226.9±0.9 and 219.5±1.4 Ma and an apparent age of 194.5±3.0 Ma. Thus, the late Triassic collision between the Yangtze and Sino-Korean palaeocontinents has been constrained.

A study on environmental aridity over northern and southern to Qinling Mountains under climate warming

Based on the data up to 1999 from hydroclimatological departments, this pape analyzes the climatic divide implications of the Qinling Mountains in regional response to the process of climate warming, due to which the grades of dryness/wetness (GDW) in 100 years show that the northern region has entered a drought period, while the southern is a humid period. In a course of ten years, the D-value of annual average air temperature over southern Shaanxi (the Hanjiang Valley) and the Central Shaanxi Plain (the Guanzhong Plain) has narrowed, i.e., the former with a slight change and the latter with rapid increase in temperature. Both regions were arid with the decrease in precipition D-value, namely the plain became warmer while the south was drier. The Qinling Mountains play a pronounced role in the climatic divide. The runoff coefficient (RC) of the Weihe River decreases synchronously with that of the Hanjiang due to climate warming. The RC of Weihe dropped from 0.2 in the 1950s to less than 0.1 in the 1990s. The Weihe Valley (the Guanzhong Plain) is practically an arid area due to shortage of water. The successive 0.5, 1.0°C temperature anomaly over China marks, perhaps, the improtant transition period in which the environment becomes more vulnerable than before. The study shows the obvious trend of environmental aridity, which is of help to the understanding of regional response to global climate change.

Comparison of the Typical Metallogenic Systems in the North Slope of the Tongbai-East Qinling Mountains and its Geologic Implications

The Tongbai-East Qinling Mountains, an important part of the Central orogenic belt, is one of the most important metallogenic belts in China and contains lots of orogenic-type and VMS-type （Volcanogenic Massive Sulfide type） metallogenic systems. The Dahe and Shuidongling VMS-type Cu-Zn deposits, located in the Erlangping Group in Tongbai and East Qinling Mountains, respectively, show similar geological and geochemical features. The Huoshenmiao Formation in the East Qinling region and the Liushanyan Formation in the Tongbai region are spilite-keratophyre sequences occurring in the western and eastern sides of the Nanyang Basin, respectively, and are interpreted to be equivalent to each other. The orogenic-type Au-Ag deposits can be subdivided into two styles; namely, fault- or structure-controlled （e.g. Yindonggou） and stratabound （e.g. Poshan）. The Poshan and Yindongpo orogenic-type Au-Ag deposits, whose ore bodies are strictly hosted in carbonaceous strata in the Tongbai Mountains, show obvious stratabound characteristics. Their ore-fluids are enriched in K^＋ and SO^2-4 and are regarded as K^＋-SO^2-4 types. The Pb-isotope ratios of sulfides of the ores are extremely uniform and significantly different from those of the tectonostratigraphic terranes of the Qinling orogens except for the ore-hosting strata of the Waitoushan Formation. The Yindonggou and Xuyaogou orogenic Au-Ag deposits in the East Qinling Mountains, whose ore bodies are hosted in the faults cutting the hosting strata or granite body, show fault-controlled characteristics. Their ore-fluids belong to the Na^＋-Cl^- type. The Pb-isotope ratios of sulfides of ores are similar to those of the northern Qinling orogenic belt. The Waitoushan Formation, dominated by carbonaceous sericite-rich schists and only occurring in Tongbai region, should be detached from the Erlangping Group, which occurs both in the western and eastern sides of the Nanyang Basin. Future ore exploration in the Tongbai-East Qinling Mountains should focus on fault-controlled Au-Ag lodes.

Structural patterns of land types and optimal allocation of land use in Qinling Mountains

The case study based on Qinling Mountains in Shaanxi Province of China, in virtue of the information from TM image, classifies the land types and analyzes their spatial and temporal differential law, and puts forward three structural patterns of land types in mountainous areas, namely, spatial, quantitative and qualitative structures of mountainous land types. Furthermore, it has been noticed that the analysis of structural patterns can disclose the heterogeneity and orderliness of combination of land types, which can lay the theoretic foundation for comprehensively recognizing ecological characteristics and succession law of structure and function of land types. After the all-around comparative analysis, an optimal allocation of land use in Qinling Mountains has been put forward according to the principle of sustainable development and landscape ecology, which can lay the scientific foundation in practice for the structural adjustment and distribution optimization from the macro level to micro level.

Effects of Environmental Factors on Tree Seedling Regeneration in a Pine-oak Mixed Forest in the Qinling Mountains, China

Natural seedling regeneration and tree establishment are affected by various environmental factors. In this study, we established eight, eight, six, and four independent forest stands （each stand was further divided into five subplots） respectively based on the altitudinal gradient, stand density, slope location, and slope aspect to investigate the effects of environmental factors on tree seedling regeneration in a pine-oak mixed forest. The results indicated that the seedling density was significantly higher at altitudes of 1,283 m to 1,665 m, whereas the sapling density did not differ with altitudes. The seedling and sapling density decreased significantly at 1,835 m. The seedling densities on the upper slopes were much higher than those on the middle and lower slopes, whereas the sapling density had no difference. The sapling density decreased southwest （20°-75°） whereas gradually from the it increased on the shady slopes to the northeast （40°）. The seedling density increased from southwest （20°） to northeast （40°The seedling and sapling densities increased with the stand density （850 trees ha-1 to 1,525 trees ha-1） whereas the sapling density was significantly lower in stands （1,9oo trees ha-O. Principal components analysis showed that the slope aspect and stand density had more important roles in tree regeneration in this study region compared with the other two factors. Therefore, our findings suggest that it will be beneficial to keep stands at a moderate density on shady slopes. Appropriate thinning of higher density stands is also expected to promote the natural regeneration of pine-oak mixed forest.

Recent Climate Warming of Central China Reflected by Temperature-sensitive Tree Growth in the Eastern Qinling Mountains and its Linkages to the Pacific and Atlantic Oceans

We have developed a 202-year tree-ring width chronology of Shensi fir(Abies chensiensis) growing in an open canopy forest at the treeline of the eastern Qinling Mountains. Climate response analyses revealed that the ring width of Shensi fir trees is primarily controlled by the range of temperature from February–June. The regression model that we used for statistical temperature reconstruction passed the leave-one-out cross-validation used in dendroclimatology, resulting in a quality-controlled February–June reconstruction for the eastern Qinling Mountains. The model accounts for 36.7% of the instrumental temperature variance during the period of 1960–2012. Warm springs and early summers occurred during AD 1870–1873, 1909–1914, 1927–1958 and 1997–2012, while the periods of AD 1874–1908, 1915–1926 and 1959–1996 were relatively cold. Spatial climate correlation analyses with gridded land surface data revealed that our temperature reconstruction contains a strong regional temperature signal for central China. The linkages of ourtemperature reconstruction with sea surface temperature in the Atlantic and Pacific oceans suggest the connection of regional temperature variations to large-scale ocean–atmosphere–land circulation. Preliminary analysis of links between large-scale climatic variation and the temperature reconstruction also shows that there is a relationship between extremes in spring and early summer temperature and anomalous atmospheric circulation in the Qinling Mountains. Overall, our study provides reliable information for the research of past temperature variability in the Qinling Mountains, China.

Stable Carbon Isotopic Composition of Black Carbon in Surface Soil as a Proxy for Reconstructing Vegetation on the Northern Slope of the Qinling Mountains

Because of the unique geographical location and important ecological effect of the Qinling Mountains, reconstruction of its vegetation and climate needs comprehensive research. We need to consider a multiple-proxy approach to gain more information on recovering the paleovegetation and climate in the Qinling Mountains. Black carbon （BC） is produced by the incomplete combustion of vegetation and fossil fuels, and is a good proxy, recording paleoenvironmental information. However, in the Qinling Mountains, what are the characteristics of the BC, and whether BC stable carbon isotope （δ^13CBc） can be used as a new proxy to study ancient vegetation, still need further study. In order to establish a sound basis for studying paleoenvironmental by BC proxy in the Qinling Mountains, we carried out systematic and detailed study on modern process of BC on the northern slope of the mountains. We analyzed stable carbon isotopes and carbon concentration of organic carbon （% SOC, δ^13Csoc） and BC （%BC, δ^13CBc）, and identified the pollen assemblages from systematically sampled surface soil. The results show that the calculated ratio of C4 plants in the vegetation （%C4） based on the δ13Csoc data reflects a similar distribution of C4 plants in the surface vegetation and the pollen assemblage. The δ^13Cac values have a strong positive correlation with δ13Csoc values, and their difference （△13CSOC-BC） is in the low range. These data indicate that δ^13CBC and δ^13CSOC have very similar characteristics. Surface soil δ13BC values can indicate surface vegetation as effectively as δ^13Csoc values, and the δ^13CBC proxy can be used effectively in paleovegetational research in the northern slope of Qinling Mountains.

Tree-ring response of Larix chinensis on regional climate and sea-surface temperature variations in alpine timberline in the Qinling Mountains

Tree-ring width chronologies of Larix chinensis were developed from the northern and southern slopes of the Qinling Mountains in Shaanxi Province,and climatic factors affecting the tree-ring widths of L.chinensis were examined.Correlation analysis showed that similar correlations between tree-ring width chronologies and climatic factors demonstrated that radial growth responded to climate change on both slopes.The radial growth of L.chinensis was mainly limited by temperature,especially the growing season.In contrast,both chronologies were negatively correlated with precipitation in May of the previous year and April of the current year.Spatial climate-correlation analyses with gridded land-surface climate data revealed that our tree-ring width chronologies contained a strong regional temperature signal over much of northcentral and eastern China.Spatial correlation with seasurface temperature fields highlights the influence of the Pacific Ocean,Indian Ocean,and North Atlantic Ocean.Wavelet coherence analysis indicated the existence of some decadal and interannual cycles in the two tree-ring width chronologies.This may suggest the influences of El Nin˜o-Southern Oscillation and solar activity on tree growth in the Qinling Mountains.These findings will help us understand the growth response of L.chinensis to climate change in the Qinling region,and they provide critical information for future climate reconstructions based on this species in semi-humid regions.

Selenium Enrichment in Cambrian Stratabound Gold Deposits in the Western Qinling Mountains

Stratabound gold deposits in the western Qinling Mountains occur in Cambrian chert formation composed of carbonaceous chert and carbonaceous slate. The distinctive chert formation provides important grounds for the mineralization and controls on the formation of gold deposits. Study shows that Se is exceptionally higher in both host rocks and gold orebodies. It may be recovered as a valuable component in ores for total utilization, and in some localities even independent Se orebodies (which are mined exclusively for Se) may be delineated. In gold ore Se mainly occurs as independent minerals or in the isomorphous form in sulphides and there is a positive correlation between Se and Au.

Fine-scale activity patterns of large-and medium-sized mammals in a deciduous broadleaf forest in the Qinling Mountains,China

The composition of animal species and interactions among them are widely known to shape ecological communities and fine-scale(e.g.,<1 km)monitoring of animal communities is essential for understanding the relationships among animals and plants.Although the co-existence of large-and medium-sized species has been studied across different scales,research on fine-scale interactions of herbivores in deciduous broadleaf forests is limited.Camera trapping of large-and medium-sized mammals was carried out over a 1 year period within a 25 ha deciduous broadleaf forest dynamics plot in the Qinling Mountains,China.Fourteen species of large-and medium-sized mammals,including six carnivores,six ungulates,one primate and one rodent species were found.Kernel density estimations were used to analyse the diel or 24 h activity patterns of all species with more than 40 independent detections and general linear models were developed to explore the spatial relationships among the species.The combination of overlapping diel activity patterns and spatial associations showed obvious niche separation among six species:giant panda(Ailuropoda melanoleuca David),takin(Budorcas taxicolor Hodgson),Reeves’s muntjac(Muntiacus reevesi Ogilby),tufted deer(Elaphodus cephalophus Milne-Edwards),Chinese serow(Capricornis milneedwardsii David)and wild boar(Sus scrofa Linnaeus).Long-term fine-scale monitoring is useful for providing information about the co-existence of species and their interactions.The results demonstrate the importance for fine-scale monitoring of animals and plants for improving understanding of species interactions and community dynamics.

The Loess in the Qinling Mountains and Its Records of Palaeoenvironmental Changes

: The Fengzhou loess section is very typical in the Qinling Mountains. The section, which is about 82 m thick and underlain by Neogene red clays, consists of 33 layers of loess and 33 layers of palaeosol. It covers the Brunhes normal polarity zone and Matuyama reversed polarity zone, and the B / M boundary is located in the middle part of loess layer 8 (L8). The loess of the Matuyama reversed polarity zone records the Jaramillo, Olduvai and Reunion normal polarity subchrons. The boundary between the Matuyama reversed polarity zone and Gauss normal polarity zone (M / Ga) appears on the lithological boundary between loess and Neogene red clays. Loess accumulation in the Fengzhou section started before 2.48 Ma B.P. Based on the stratigraphical structure, the material composition and magnetic susceptibility curve of the Fengzhou loess section, the palaeoclimatic changes during the last 2.48 Ma in the Qinling Mountains are subdivided into 66 cold-dry and warm-humid stages, equivalent to 33 climatic cycles. The above climatic fluctuations agree very well with those of the oxygen isotopic records from deep—sea deposits.

Study on Dominant Expression of Regional Culture of the Qinling Mountains in Local Residential Buildings

Regional culture of the Qinling Mountains shows distinct features since it was born in the local outstanding ecological environment, study on local architecture is significant for the dominant expression of regional culture, protection of local environment, and echoing with the theme of ecological civilization construction. This paper, on the basis of the mutual infl uence and evolution of regional culture and style of local residential buildings, explored the reasons for the weakening of local architectural style, and specified the signifi cance of promoting local style of the living environment. By studying the infl uence of local natural environment and humanistic environment on architectural style along the northern foot of the Qinling Mountains, the paper explored the expression of regional culture in residential buildings, with Xian Garden(Xi'an Yuanzi) as an example, and aimed at giving useful help to the dominant expression of regional culture in modern residential buildings.

Cosmogenic ^(26)Al/^(10)Be Burial Dating of the Uplift Rate of Southern Qinling Mountains since the Middle Pleistocene

Objective The Qinling Mountains （QM） in Central China is a natural harrier that corresponds to the boundary between the southern and northern climate and environment （Gong Hujun et al., 2017）. Northern QM is relatively steep, and southern QM is in contrast relatively low and gentle. Investigations have shown that the average uplift rate of northern QM since 17.8 Ma is approximately 0.19 mm/a （Yin Gongming et al., 2001）, whereas that of central QM since 0.36 Ma is approximately 0.32 mm/a （Wang Fei et al., 2004）. To date, however, few investigations have yet been conducted on the uplift rate of southern QM. Accordingly, we aim to obtain the uplift rate of southern QM by using the cosmogenic ^26A1/^10 Be burial dating method to determine the age of the highest river terrace on the southern slope of QM.

Runoff projections of the Qinling Mountains and their impact on water demand of Guanzhong region in Northwest China

The Qinling Mountains are the main source of water supply to the Weihe River and Hanjiang River,but systematic studies on runoff changes in this region are lacking.This study simulated the annual runoff in the Qinling Mountains from 1970 to 2015 using the Variable Infiltration Capacity(VIC)hydrological model and projected runoff changes from 2015 to 2100 under different Shared Socio-economic Pathways(SSPs).Moreover,the water demand in the north and south of the Qinling Mountains from 2019 to 2100 was analyzed by constructing a system dynamic method.The results show that the mean annual runoff in the southern and northern slopes of the Qinling Mountains from 1970 to 2015 were 21.5×10^(9) and 3.52×10^(9) m^(3),respectively.Under the SSP1-2.6,SSP2-4.5,SSP3-7.0,and SSP5-8.5 scenarios from 2015 to 2100,the mean annual runoff in the southern(northern)slope will increase by 2.9(1.5),2.0(1.4),2.7(1.6),and-2.1(0.4)×10^(9) m^(3),respectively.The mean annual water consumption in the northern foothills of the Qinling Mountains for Baoji,Xi’an,and Weinan cities is projected to increase from 4.1×10^(9) m^(3) in 2015 to 7.6×10^(9) m^(3) in 2100,while that in the southern foothills for Hanzhong,Ankang,and Shangluo cites is projected to increase from 2.7×10^(9) m^(3) in 2015 to 4.8×10^(9) m^(3) in 2100.Thus,runoff in the southern slope is much larger than the water demand in the southern foothill.The increase of runoff in the northern slope cannot satisfy the water demand in the northern foothill.Therefore,the South-to-North Water Transfer project is proposed for meeting the water demand in the northern foothill.Under the premise of ensuring downstream water demand,approximately 10.5×10^(9) m^(3) of water resources can be transferred from the southern slope to the Weihe River and the Beijing-Tianjin-Hebei-Henan regions under SSP1-2.6,but only 3.9×10^(9) m^(3) can be transferred under SSP5-8.5.

The features of physical geography in the transitional region between Qinling Mountains and Huanghuai Plain

The features of physical geography in the transitional region between Qinling Mountains and Huanghuai Plain possess transitional characters evidently in two directions: one is from the western mountain to the eastern plain and the other is from southern subtropical zone to northern temperate zone. Torrential rain, especially strong torrential rain is frequent in the transitional region, and there are many torrential rain centers. A majority of torrential rain is distributed among 100-200 m asl. The winter temperature at 100-400 m asl is higher than that in Huanghuai Plain whose altitude is lower than that of the transitional region, and the highest temperature in January appears at 350-400 m asl.The thickness of warm slope belt in the transitional region varies from 100 m to 250 m asl. The formation of torrential rain and warm slope belt is the result of joint action of atmospheric circulation and local terrain. Frequent torrential rains and warm slope belt had tremendous influences on the soil properties, plant distribution and local climate in the transitional region.

Ecological variations of woody species along an altitudinal gradient in the Qinling Mountains of Central China:area-based versus mass-based expression of leaf traits

Leaf trait-based research has become the preferred method to understand the ecological strategies of plants.However,there is still a debate on whether area-based or mass-based traits provide different insights into environmental adaptations and responses.In this study,seven key leaf traits(maximum net photosynthetic rate,dark respiration rate,nitrogen content,photosynthetic nitrogen use efficiency,leaf mass per area,leaf dry matter contents and leaf area)of 43 woody species were quantified on the basis of both area and mass along an altitudinal gradient(1100–2700 m)in the Qinling Mountains of China.Differences in leaf traits and bivariate correlations between the two expressions were compared.By considering different expressions,the strengths and directions of the responses of leaf traits to the altitudinal gradient were determined.Leaf traits showed large variations;interspecific variations contributed more to total variance than intraspecific variations.Bivariate correlations between photosynthetic traits and structural traits(mass per area,dry matter content,and area)were weaker on a mass basis than those on an area basis.Most traits exhibited quadratic trends along the altitudinal gradient,and these patterns were more noticeable for area-based than mass-based traits.Area-based traits were more sensitive to changes in temperature and precipitation associated with altitude.These results provide evidence that mass-versus area-based traits show different ecological responses to environmental conditions associated with altitude,even if they do not contain very broad spatial scales.Our results also indicate distinction of photosynthetic acclimation among the two expressions along an altitudinal gradient,reflecting trade-offs among leaf structure and physiological traits.