Evolutionary Significance of Pylentonemid Radiolarians and Their Late Devonian Species from Southwestern Tianshan,China

A group of radiolarian fossils with a complete morphological gradient of increasing polarization are reported from upper Devonian successions in southwestern Tianshan, China. Specifically, the important transitional mophotypes, Pylentonema transitorum sp. nov., Archocyrtium medium sp. nov. and Quadrapesus transitivus sp. nov, were found. Based on the fact that the new transitional radiolarian fossils resemble more closely to pylentonemids （cyrtoid nassellarians） than to entactinids （spumellarians） in morphology, they are assigned to pylentonemids. Pylentonemids share characters of both the Nassellaria （e.g., horned cephalis, basal opening with pylome） and Spumellaria （e.g., system of trabecular spicules）, and they are thus of evolutionary significance. The evolutionary trend from spumellarians to nassellarians is characterized by （1） internal spicule： from simple trabecular spicules to complicate components （A;Ir,II;D;V;Lr,Li;MB）; （2） polarization of the radiolarian tests due to rearrangment of spine, from radial to an axial symmetry; （3） the gradual formation of a basal opening （aperture, pylome, or podome）; and the transition from a pseudopylome, a primative pylome with a narrow margin on the outer shell, to a true pylome rimmed with a narrow, elevated impermite wall.

Discovery of Radiolarian Fossils from the Aiketik Group at the Western End of the South Tianshan Mountains of Chinaand Its Implications

The Aiketik Group, distributed at the western end of the South TianshanMountains, China, is an important lithostratigraphic unit involved in the South Tianshan orogen. Itis separated from the adjacent rocks by faults. Generally, the geologists ascribed it to the UpperCarboniferous according to Pseudostaffella sp., Profusulinella sp. and Fusulinella sp. found fromthe limestone and sandy limestone of Aiketik. Our radiolarian fossils were obtained from the chertsamples collected from the Haladaok section located at the upper Tuoshihan River. The fossils mainlyinclude Albaillella undulata Deflandre, Albaillella paradoxa Deflandre, Albaillella sp. aff. A.paradoxa Deflandre, Albaillella sp. cf. A. deflandrei Gourmelon, Albaillella sp., Albaillellaexcelsa Ishiga, Kito and Imoto (?), Belowea variabilis (Ormiston et Lane), Callella cf. C.parvispinosa Won, Entactinia cf. E. tortispina Ormiston et Lane, Entactinia aff. E. tortispinaOrmiston et Lane, Entactinia variospina Won, Entactinia sp., Eostylodictya rota (Won),Latentifistula impella (Ormistone et Lane) (?), Latentifistula turgita Omiston et Lane,Latentifistulidae gen. et. sp. indet. and Polyentactinia cf. aranea Gourmelon. Among them,Albaillella excelsa Ishiga, Kito and Imoto (?) is a Late Permian species with some elementsuncertain as there is only one poorly-preserved fossil of this species found so far. And tworadiolarian assemblages can be identified from the other fossils. One is the early EarlyCarboniferous assemblage represented by Albaillella undulata Deflandre, Albaillella paradoxa andAlbaillella sp. cf. A. deflandrei Gourmelon. And the other is the late Early Carboniferousassemblage represented by Eostylodictya rota (Won). This is the first discovery of radiolarianfossils in the Aiketik Group, also the first discovery of Late Permian radiolarian fossils in theSouth Tianshan Mountains. Meanwhile, this is the current westernmost sampling site of radiolarianfossils in the South Tianshan Mountains.

Middle Jurassic-early Cretaceous radiolarian assemblages of the western Yarlung Zangbo Suture Zone: Implications for the evolution of the Neo-Tethys

Cherts in the Zhongba melange of the western Yarlung Zangbo Suture Zone(YZSZ) contain well preserved radiolarian assemblages. These radiolarian assemblages indicate that the Zhongba melange has middle Jurassic-early Cretaceous remnant, are coeval with those from the central and eastern parts of the YZSZ. Cherts from the Najiu area yield Aalenian to Aptian radiolarians, while cherts interbedded with siliceous mudstones from the Bielongjiala area yield Aptian radiolarians, indicating that terrigenousderived sediments were deposited during early Aptian. The above observations indicate that the entire YZSZ have a similar geochronological framework and thus they underwent similar geological evolution:(1) during the Jurassic, the Neo-Tethys was a wide ocean with pelagic sediments distal from continents;(2) during the Cretaceous(around 130-120 Ma), the Neo-Tethys started to subduct along the southern margin of the Lhasa block, and terrigenous-derived siliceous mudstone began deposition.

Early Eocene Radiolarian Fauna from the Sangdanlin, Southern Tibet: Constraints on the Timing of Initial India-Asia Collision

This is a new report on the early Eocene radiolarian fauna from the Sangdanlin section in the Gyirong region, along the southern margin of the Yarlung Zangbo Suture Zone. The Sangdanlin section measured in this study is divided into three lithostratigraphic units from bottom to top： the Zongzhuo, Sangdanlin, and Zheya formations. Abundant radiolarian fossils were obtained from the Sangdanlin section and 54 species of 30 genera were identified and assigned as follows： Cryptamphorella conara-C. macropora the late Cretaceous Zone and Amphis_phaera coronate, Bur）ella tetradica-Bekoma campechensis, and B.bidartensis-B. divaricata the Paleocene-early Eocene Interval Zones. The Paleocene- early Eocene radiolarian zones are comparable to the radiolarian zones RP4-RP8 in New Zealand. Based on the data of radiolaria and lithofacies, it is suggested that the Zongzhuo Formation should be deposited along the base of the north-facing, continental slope of the Greater Indian continental margin, and the Sangdanlin Formation should be a deep marine, sedimentary sequence located in a foreland basin. The early Eocene radiolarian fauna in the Sangdanlin Formation constrains the initial age of the India-Asia collision to no later than 53.6 Ma.

Geochronology and Geochemistry of the Radiolarian Cherts of the Mada'er Area,Southwestern Tianshan:Implications for Depositional Environment

In the southwestern Tianshan,the geologic ages of many strata and their depositional environments are still poorly constrained because of complex structures.The Mada＇er area is located in the Kuokesaleling belt,Southwestern Tianshan.The cherts from the former Wupata＇erkan Group contain abundant radiolarian fossils,including 10 species which are identified as late Devonian to early Carboniferous in age.Eleven chert samples have SiO_2 contents ranging from 88.80 wt%to 93.28 wt%, and 2.02 wt%to 3.72 wt%for Al_2O_3.The SiO_2/Al_2O_3 ratios of all samples vary from 23.84 to 46.11, much lower than those of the pure cherts（80-1400）.These values suggest that the cherts contain high ratios of terrigenous materials.The Al_2O_3/（Al_2O_3＋Fe_2O_3）ratios vary between 0.64 and 0.77,whereas V and Cu concentrations range from 10.92 ppm to 26.7 ppm and from 2.15 ppm to 34.1 ppm respectively. The Ti/V ratios vary from 25.53 to 44.93.The total REE concentrations of the cherts are between 30.78 ppm and 59.26 ppm,averaging 45.46 ppm.The（La/Ce）_N ratios range from 0.81 to 1.12,and 0.88-1.33 for（La/Yb）_N,averaging 1.09,which suggests a continental margin environment.Consequently,it is inferred that the cherts formed in a residual sea environment during the late Devonian to early Carboniferous time,which suggests that the collision between the Karakum-Tarim and Kazakhstan-Junggar plates did not occur at the time.In addition,the regional geological information indicates that the study area experienced a post-collision stage during the early Permian,and thus it is likely that the collision between the two plates took place in the late Carboniferous.

Reexamination of Radiolarian Biostratigraphy in Permian in Pelagic Chert Sequences at Dachongling Section, South China

Permian radiolarian biostratigraphy was reexamined in the bedded chert section in Qinzhou area of southeast Guangxi Zhuang Autonomous Region, South China. On the basis of the analyses of characteristic radiolarian composition, six radiolarian zones are recognized, namely, Pseudoalbaillella longtanensis Zone, Pseudoalbaillella globosa Zone, Follicucullus monacanthus Zone, Follicucullus scholasticus Zone, Follicucullus charveti Zone and Neoalbaillella ornithoformis Zone, in ascending order. Correlation of these radiolarian zones with those in Japan is presented. The Neoalbaillella ornithoformis Zone was firstly discovered in Dachongling Section, which may indicate that pelagic chert of the late Permian existed there, according to the co occurring conodonts.

Abrupt changes of radiolarian fauna at 600 and 120 ka B. P. in the southern South China Sea and their paleoceanographic implications

Abrupt changes in radiolarian composition are discovered over the last 600 and 120 ka B. P. based on quantitative analyses of radiolarians in ~ 17957 - 2 of the southern South China Sea. The distinct changes at 600 ka B. P. could correspond to the onset of the 100 ka cycle during the glacial and interglacial periods. This abrupt change in the 100 ka cyclicity at 600 ka B. P. occurred also in the magnetic susceptibility signal that is obtained from and paleosol sequences of the China Loss Plateau. The larger amplitude and stronger cyclicity in the susceptibility signal after 600 ka B. P. reflect the prominent change in the intensity of the monsoon, induced by an enhancement of the momsoon circula- tion. Stronger seasonality during the glacial period in the South China Sea, resulted from strengthening of winter monsoon, might lead to the changes in the radiolarian composition at 600 and 120 ka B. P. It can be suggested that only species adapted to a broader temperature range might have been able to live in this environment. Therefore, the abrupt changes in radiolarian composition at 600 and 120 ka B. P. could be attributed to the stronger so differences between summer and winter that were caused by the striking change in the intensity of the monsoon circulation.

Preservation of Radiolarian Diversity and Abundance in Surface Sediments of the South China Sea and Its Environmental Implication

Quantitative analysis of radiolarian preservation in surface sediments from different regional environments of the South China Sea （SCS） is discussed in this article for illustrating their relationship. The results show that species diversity and abundance of radiolarian in the surface sediment increase with water depth, indicating no obvious dissolution of radiolarian shell in the deep SCS. Upwelling activity and volcanic eruption are both in favor of radiolarian propagation, and cause the settlement enrichment of radiolarian shell. Whereas, an abnormal reduced radiolarian fauna in deeper slope sediment may result from the remain of turbid flow that brings sediments from shallow area, such as shelf and upper slope. Radiolarian species are mainly composed of tropic-subtropic types, and include some cold or polar species, which indicates that a mistake in paleoceanographic analysis should arise from the monospecific index. However, synthesis radiolarian indexes in sediments may well respond to the ecologic and depositional environments in the SCS.

Characteristics of Albaillella (Albaillellarian,Radiolarian) Fauna from Guadalupian to Lopingian Series in Permian,South China

On the basis of establishment of radiolarian biostratigraphy and conodont biostratigraphy, a radiolarian Albaillella fauna in the transitional environment from Guadalupian to Lopingian Series in Permian was found at a pelagic chert section in southeast Guangxi, South China. Radiolarian Albaillella is one of the most sensitive biology to the transitional environment. The Albaillella fauna shows an ecological evolutionary process from Guadalupian to Lopingian: declined stage-recovery stage-flourishing stage. The study of characteristics of the Albaillella fauna in the transitional environment may provide more information, not only for the subdivision and correlation of a high-resolution biostratigraphy, but also for influence of radiolarian Albaillella fauna on the pre-Lopingian mass extinction.

Late Olenekian Radiolarians from Bedded Chert of Ashio Terrane, Northeast Japan, and Faunal Turnovers in Western Panthalassa during Early Triassic

Late Olenekian assemblages in the western Panthalassa have been recovered from bedded radiolarian chert sequences of an accretionary complex, the Ashio belt. These faunas are documented and considered in terms of radiolarian diversity and faunal turnover during the latest Permian to Middle Triassic time. The fauna includes 30 radiolarians belonging to Spumellaria or Entactinaria, with two relicts from the Late Permian. This late Olenekian fauna is markedly different from Permian and Anisian faunas, respectively, and is herein named the Minowa fauna. Study of the literature indicates that radiolarian provinces were significantly disconnected between the western Panthalassa and eastern Tethys during late Olenekian time. Furthermore, 121 of 143 species disappeared during late Olenekian time, and in turn 118 new species appeared in the western Panthalassa around the Olenekian Anisian boundary. It is concluded that faunal turnover occurred at least three times between the latest Permian and Middle Triassic. The first turnover is the Paleozoic type radiolarian extinction at the Permian Triassic boundary, the second is the diversification of spheroidal Spumellaria and Entactinaria between early and late Olenekian time, and the third is a faunal turnover from the Minowa fauna to the true Mesozoic type radiolarian faunas that are characterized by multi segmented Nassellaria.

Late Jurassic and Early Cretaceous Radiolarians from Zhilong,Gyangse,Tibet

Abundant well-preserved Late Jurassic-Early Cretaceous radiolarian fossils were extracted from 31 bedded green chert samples collected from Zhilong section,Gyangse,southern Tibet.This

Radiolarians from the Pohang Basin,Southeast Korea and Paleoceanographic Implications

We identified a total of 101 species and two subspecies of radiolarians belonging to 56 genera from 95 samples collected from the Hagjeon and Duho Formations in the Pohang Basin of the southeastern Korean Peninsula. On the basis of the biostratigraphic range of Cyrtocapsella cornuta and Theocorys redondoensis, the depositional period of the upper Hagjeon and lowest Duho Formations was determined to be early to late Middle Miocene. The occurrence of deep-dwelling radiolarians indicates that the paleobathymetry seems to become gradually progressing toward an upper bathyal environment in the middle part of the Hagjeon Formation. However, we prefer to accept another interpretation for the occurrence of deep-sea indicators in the Hagjeon and the lowest part of the Duho Formations, and consider the presence of a region of upwelling cold water that might have simulated a deep-water environment in relatively shallow water. This interpretation is based on the present upwelling of a cold-water mass off the southeast coast of Korea, the occurrence of upwelling microfossils from the Pohang Basin, and the effect of the closing of the Korea Strait approximately 15 Ma. We also considered that the uppermost part of the studied section represents a shallow-water environment.

New Discovery of Radiolarians in the Pomulong Mélange, Middle part of the Yarlung Zangbo Suture Zone, Tibet

Objective The Pomulong mélange was established in the regional geological survey of Lhaze County at the scale of 1:250000 by the Geological Survey Institute of Hubei Province.

First Report of Redeposited Cretaceous Radiolarians in the Eocene Sand-Shale Member of Zhepure Formation, Tüna, Yadong, Tibet

Cretaceous radiolarians were firstly reported from the Eocene Sand-Shale Member of Zhepure Formation in the Gulupu section, Tüna, Yadong, Tibet. In spite of poor preservation, 24 species of 14 radiolarian genera have still been identified, clearly indicating an age, ranging from early Aptian–Maastrichtian, and therefore should have occurred in the Eocene as a result of redeposition. The study of the stratigraphic origin of these radiolarians will shed light on the paleoenvironment and late evolutionary history of the Tibet-Tethys.

Micropalaeontological and Geochemical Evidence of the Late Jurassic Radiolarians Cherts of Naga Ophiolite Hill, Nagaland, Northeast-India

We analyzed the radiolarian assemblages (pelagic sediments) and geochemical studies of 15 samples collected from Meluri district Nagaland, Naga Hills Ophiolite belt (NHO) and it consists of the Flysch Naga-Arakan-Yoma trough beyond the Indo-Myanmar border. It reflects one of the many components of the Himalayan Orogenic systems of Tethyan oceanic crust discovered in Nagaland at Meluri district of India and they have very well-preserved studies and can be identified to Upper Jurassic (Kimmeridgian-mid-Tithonian). As well as a continuous sequence of Kimmeridgian-mid-Tithonian, five radiolarian assemblages have been identified in this study. This is so far the first recovery of pelagic sediment assemblages (Radiolarians) from the Naga Hills Ophiolite complex. Samples of Naga Hills Ophiolite belts fell in the Sublitharenite, Litharenite, and Fe-shale Fe-sand area, and the A-CN-K diagram indicates that these sediments in the source region were exposed deep to moderate weathering conditions. In the discriminant plot, the Naga Hill-Ophiolite-Chert samples occupy the field of origin of the sediment quartzose, and the log (K2O/Na2O) versus SiO2 samples fall into the active continental marginal field, and only one sample falls into an island arch, suggesting that the tectonic framework of the Naga Hills-Chert samples was deposited in the active continental margin and the diagrams K2O/Al2O3 and MgO/Al2O3 illustrated the tectonic setting of the marine environment from Naga Hill Ophiolite Chert. Such findings have been consistent with the current geology of Naga Hills in the province of Nagaland.

Paleocene deep-water sediments and radiolarian faunas: Implications for evolution of Yarlung-Zangbo foreland basin, southern Tibet

This is the first report on the Paleocene deep-water sequences and radiolarian faunas, which are distributed along the southern side of the Yarlung-Zangbo suture zone. The Zheba group is coined to indicate these Paleocene sequences which are subsequently divided into two lithostratigraphic units based on the lithology observed in the field. The lower unit characterized by the rhythmic cherts and siliceous shales is named the Sangdanlin formation, and the upper one composed mainly of flysches is termed the Zheya formation. The radiolarian faunas from the Zheba group are assigned to the RP1-RP6 zones of the Paleocene age. The Early Paleocene ra- diolarian assemblages have the potential to be established into the low latitude radiolarian zones and to fill in the gap between the Late Cretaceous and the Late Paleocene radiolarian zonations. The radiolarian dating provides a valuable tool for the regional correlation and reconstruction of the sedimentary environment of the Neo-Tethyan Ocean. The preliminary work shows that the Paleo- cene sequences accumulated in a foreland basin resulted from the southern Asian margin loading onto the northern Indian passive continental margin. The Yarlung-Zangbo foreland basin se- quences deposited on the Indian passive continental margin also resulted in many good source- reservoir-covering assemblages for oil and gas resources.

Mesozoic radiolarian chert from the middle sector of the Yarlung Zangbo suture zone,Tibet and its tectonic implications

Three radiolarian fauna, aged as the Middle-Late Triassic, Late Jurassic-Early Creta- ceous and Cretaceous radiolarian fauna, have been recognised in the radiolarian cherts from the middle sector of the Yarlung Zangbo suture zone, southern Tibet, China. The average contents of SiO2 in the radiolarian cherts of the Middle-Late Triassic and Late Jurassic-Early Cretaceous are 90.24% and 92.58% respectively, with average ratios of Al/(Al+Fe+Mn) as 0.75 and 0.74 respectively, the average ratios of MnO/TiO2 as 0.36 and 1.24, the average ratios of Ce/Ce* as 1.15 and 1.03, and the average ratios of LaN/CeN as 0.85 and 0.93. These geochemical features indicate that both of them are biogenic, deposited in a continental margin basin. The SiO2 content of the Early Cretaceous radiolarian chert is 94.12%, with the ratio of Al/(Al+Fe+Mn) as 0.59, ratio of MnO/TiO2 as 4.30, ratio of Ce/Ce* as 0.60, ratio of LaN/CeN as 1.59, which imply that the chert is biogenic and was deposited in a pelagic basin. The Middle-Late Triassic association of the radiolarian chert and turbidites as well as their geochemical characteristics indicates the existence of a strong rifting marginal basin in the belt of the Yarlung Zangbo River then. The association of radiolarian chert and bedded basalt indicate an initial Tethyan ocean basin in southern Tibet during the Late Jurassic-Early Cretaceous. The early Cretaceous radiolarian chert coexisting with pillow basalt in top of the ophiolite suite represents sediments from the oceanic Tethyan basin.

Application of radiolarians and other fossils in non-Smith strata——Exemplified by the A'nyemaqen melange belt in East Kunlun Mts.

The main character of melange strata in an erogenic belt is the integration of mixed materials due to the superposition, displacement or loss of various tectonic slices (blocks) of different origins and environments, different scales, different grades of deformation and metamor-phism, and different stages of tectonic evolution. The approach to non-Smith strata in an erogenic belt is to focus on the understanding of the age, facies, tectonic setting of the original formation and the process of deformation-metamorphism of each tectonic slice, reconstruct the history of dispersal and integration of these tectonic slices in time and space, i.e. 4-dimensional. This paper studies the age and facies of the original formation of tectonic slices in the A’nyemaqen melange belt based on new data of radiolarians, sporo-pollen and trace fossils, and cast new lights on the research of the evolution process of that belt.

Pleistocene radiolarian biostratigraphy in the South China Sea

Based on a quantitative radiolarian analysis, 5 radiolarian zones (NR1-NR5) have been defined at Core 17957-2 from the South China Sea, considering radiolarian zonations from low latitudes. The absolute age assignment of the radiolarian zones and the ranges of the six marker species is based on the direct correlation with the paleomagnetic and isotopic record, respectively. A comparison of the radiolarian stratigraphic data obtained from Core 17957-2 from the South China Sea with those from the equatorial Pacific and the tropical Indian Ocean shows a close similarity to the ages defined in the equatorial Pacific. The obtained biostratigraphic data provide an excellent tool for further dating of Pleistocene sediments in the China Sea.