Enhanced and asymmetric signatures of hybridization at climatic margins: Evidence from closely related dioecious fig species

Hybridization plays a significant role in biological evolution. However, it is not clear whether ecological contingency differentially influences likelihood of hybridization, particularly at ecological margins where parental species may exhibit reduced fitnesses. Moreover, it is unknown whether future ecosystem change will increase the prevalence of hybridization. Ficus heterostyla and F. squamosa are closely related species co-distributed from southern Thailand to southwest China where hybridization, yielding viable seeds, has been documented. As a robust test of ecological factors driving hybridization, we investigated spatial hybridization signatures based on nuclear microsatellites from extensive population sampling across a widespread contact range. Both species showed high population differentiation and strong patterns of isolation by distance. Admixture estimates exposed asymmetric interspecific gene flow.Signatures of hybridization increase significantly towards higher latitude zones, peaking at the northern climatic margins. Geographic variation in reproductive phenology combined with ecologically challenging marginal habitats may promote this phenomenon. Our work is a first systematic evaluation of such patterns in a comprehensive, latitudinally-based clinal context, and indicates that tendency to hybridize appears strongly influenced by environmental conditions. Moreover, that future climate change scenarios will likely alter and possibly augment cases of hybridization at ecosystem scales.

Mobile bed thickness in skewed asymmetric oscillatory sheet flows

A new instantaneous mobile bed thickness model is presented for sediment transport in skewed asymmetric oscillatory sheet flows. The proposed model includes a basic bed load part and a suspended load part related to the Shields parameter, and takes into account the effects of mass conservation, phase-lag, and asymmetric boundary layer development, which are important in skewed asymmetric flows but usually absent in classical models. The proposed model is validated by erosion depth and sheet flow layer thickness data in both steady and unsteady flows, and applied to a new instantaneous sediment transport rate formula. With higher accuracy than classical empirical models in steady flows, the new formula can also be used for instantaneous sediment transport rate prediction in skewed asymmetric oscillatory sheet flows.

EXPERIMENTAL STUDY ON TURBULENT FEATURES IN THE NEGATIVE TRANSPORT REGION OF ASYMMETRIC PLANE CHANNEL FLOW

Turbulent features of streamwise and vertical components of velocity in the negative transport region of asymmetric plane channel flow have been studied experimentally in details. Experiments show that turbulent fluctuations in negative transport region are suppressed, and their probability distributions are far from Gaussian. Besides, the skewness factors attain their negative maxima at the position of the maximum mean velocity, whereas the flatness factors attain their positive maxima at the same position.

Homotopy analysis solutions for the asymmetric laminar flow in a porous channel with expanding or contracting walls

In this paper, the asymmetric laminar flow in a porous channel with expanding or contracting walls is investigated. The governing equations are reduced to ordinary ones by using suitable similar transformations. Homotopy analysis method （HAM） is employed to obtain the expres- sions for velocity fields. Graphs are sketched for values of parameters and associated dynamic characteristics, especially the expansion ratio, are analyzed in detail.

Numerical investigations of asymmetric flow of a micropolar fluid between two porous disks

Numerical solution is presented for the two- dimensional flow of a micropolar fluid between two porous coaxial disks of different permeability for a range of Reynolds number Re （-300≤ Re 〈 0） and permeability parameter A （1.0≤A ≤2.0）. The main flow is superimposed by the injection at the surfaces of the two disks. Von Karman＇s similarity transformations are used to reduce the governing equations of motion to a set of non-linear coupled ordinary differential equations （ODEs） in dimensionless form. An algorithm based on the finite difference method is employed to solve these ODEs and Richardson＇s extrapolation is used to obtain higher order accuracy. The results indicate that the parameters Re and A have a strong influence on the velocity and microrotation profiles, shear stresses at the disks and the position of the viscous/shear layer. The micropolar material constants cl, c2, c3 have profound effect on microrotation as compared to their effect on streamwise and axial velocity profiles. The results of micropolar fluids are compared with the results for Newtonian fluids.

A numerical study for boundary layer current and sheet flow transport induced by a skewed asymmetric wave

An analytical model with essential parameters given by a two-phase numerical model is utilized to study the net boundary layer current and sediment transport under skewed asymmetric oscillatory sheet flows. The analytical model is the first instantaneous type model that can consider phase-lag and asymmetric boundary layer development. The two-phase model supplies the essential phase-lead, instantaneous erosion depth and boundary layer development for the analytical model to enhance the understanding of velocity skewness and acceleration skewness in sediment flux and transport rate. The sediment transport difference between onshore and offshore stages caused by velocity skewness or acceleration skewness is shown to illustrate the determination of net sediment transport by the analytical model. In previous studies about sediment transport in skewed asymmetric sheet flows, the generation of net sediment transport is mainly concluded to the phase-lag effect.However, the phase-lag effect is shown important but not enough for the net sediment transport, while the skewed asymmetric boundary layer development generated net boundary layer current and mobile bed effect are key important in the transport process.

Asymptotic solutions for the asymmetric flow in a channel with porous retractable walls under a transverse magnetic field

The self-similarity solutions of the Navier-Stokes equations are constructed for an incompressible laminar flow through a uniformly porous channel with retractable walls under a transverse magnetic field. The flow is driven by the expanding or contracting walls with different permeability. The velocities of the asymmetric flow at the upper and lower walls are different in not only the magnitude but also the direction. The asymptotic solutions are well constructed with the method of boundary layer correction in two cases with large Reynolds numbers, i.e., both walls of the channel are with suction, and one of the walls is with injection while the other one is with suction. For small Reynolds number cases, the double perturbation method is used to construct the asymptotic solution. All the asymptotic results are finally verified by numerical results.

Bifurcation phenomena and control for magnetohydrodynamic flows in a smooth expanded channel

This work reports the effects of magnetic field on an electrically conducting fluid with low electrical conductivity flowing in a smooth expanded channel. The governing nonlinear magnetohydrodynamic （MHD） equations in induction- free situations are derived in the framework of MHD approximations and solved numerically using the finite-difference technique. The critical values of Reynolds number （based on upstream mean velocity and channel height） for symmetry breaking bifurcation for a sudden expansion channel （1：4） is about 36, whereas the value in the case of the smooth expansion geometry used in this work is obtained as 298, approximately （non-magnetic case）. The flow of an electrically conducting fluid in the presence of an externally applied constant magnetic field perpendicular to the plane of the flow is reduced significantly depending on the magnetic parameter （M）. It is expansion （1：4） is about 475 for the magnetic parameter M found that the critical value of Reynolds number for smooth = 2. The separating regions developed behind the smooth symmetric expansion are decreased in length for increasing values of the magnetic parameter. The bifurcation diagram is shown for a symmetric smoothly expanding channel. It is noted that the critical values of Reynolds number increase with increasing magnetic field strength.

NUMERICAL INVESTIGATION OF THE MECHANISM OF ASYMMETRIC VORTEX FLOWS

A robust iterative method suitable for the numerical simulation of high angle-of-attack vortex flows is established based upon the multiple line-vortex model(MLVM).With symmetric or asymmetric positions of sep- aration lines given,the first converged solution at an angle of attack as high as 60 degree is obtained by means of the present method.Numerical experiments for a tangent-ogive forebody indicate the viscous onset mechanism of asymmetric vortex flows over a body of revolution at high angles of attack and zero sideslip.

Evolutionary history of a desert perennial Arnebia szechenyi(Boraginaceae):Intraspecific divergence,regional expansion and asymmetric gene flow

The complex interactions of historical,geological and climatic events on plant evolution have been an important research focus for many years.However,the role of desert formation and expansion in shaping the genetic structures and demographic histories of plants occurring in arid areas has not been well explored.In the present study,we investigated the phylogeography of Arnebia szechenyi,a desert herb showing a near-circular distribution surrounding the Tengger Desert in Northwest China.We measured genetic diversity of populations using three maternally inherited chloroplast DNA(cpDNA)fragments and seven bi-paternally inherited nuclear DNA(nDNA)loci that were sequenced from individuals collected from 16 natural populations across its range and modelled current and historical potential habitats of the species.Our data indicated a considerably high level of genetic variation within A.szechenyi and noteworthy asymmetry in historical migration from the east to the west.Moreover,two nuclear genetic groups of populations were revealed,corresponding to the two geographic regions separated by the Tengger Desert.However,analysis of cpDNA data did not show significant geographic structure.The most plausible explanation for the discrepancy between our findings based on cpDNA and nDNA data is that A.szechenyi populations experienced long periods of geographic isolation followed by range expansion,which would have promoted generalized recombination of the nuclear genome.Our findings further highlight the important role that the Tengger Desert,together with the Helan Mountains,has played in the evolution of desert plants and the preservation of biodiversity in arid Northwest China.

Vortex Rossby Waves in Asymmetric Basic Flow of Typhoons

Wave ray theory is employed to study features of propagation pathways（rays） of vortex Rossby waves in typhoons with asymmetric basic flow, where the tangential asymmetric basic flow is constructed by superimposing the wavenumber-1 perturbation flow on the symmetric basic flow, and the radial basic flow is derived from the non-divergence equation. Results show that, in a certain distance, the influences of the asymmetry in the basic flow on group velocities and slopes of rays of vortex Rossby waves are mainly concentrated near the radius of maximum wind（RMW）, whereas it decreases outside the RMW. The distributions of radial and tangential group velocities of the vortex Rossby waves in the asymmetric basic flow are closely related to the azimuth location of the maximum speed of the asymmetric basic flow, and the importance of radial and tangential basic flow on the group velocities would change with radius. In addition, the stronger asymmetry in the basic flow always corresponds to faster outward energy propagation of vortex Rossby waves. In short, the group velocities, and thereby the wave energy propagation and vortex Rossby wave ray slope in typhoons, would be changed by the asymmetry of the basic flow.

Analytical and Computational Analysis of Flow Splitting in Multiple, Parallel Channels Systems

Previous analytical results on flow splitting are generalized to consider multiple boiling channels systems. The analysis is consistent with the approximations usually adopted in the use of systems codes (like RELAP5 and TRACE5, among others) commonly applied to perform safety analyses of nuclear power plants. The problem is related to multiple, identical, parallel boiling channels, connected through common plena. A theoretical model limited in scope explains this flow splitting without reversal. The unified analysis performed and the confirmatory computational results found are summarized in this paper. New maps showing the zones where this behavior is predicted are also shown considering again twin pipes. Multiple pipe systems have been found not easily amenable for analytical analysis when dealing with more than four parallel pipes. However, the particular splitting found (flow along N pipes dividing in one standalone pipe flow plus N -1 identical pipe flows) has been verified up to fourteen pipes, involving calculations in systems with even and odd number of pipes using the RELAP5 systems thermal-hydraulics code.

Effects of tip perturbation and wing locations on rolling oscillation induced by forebody vortices

The wing rock motion is frequently suffered by a wing-body configuration with low swept wing at high angle of attack. It is found from our experimental study that the tip perturbation and wing longitudinal locations affect significantly the wing rock motion of a wing-body. The natural tip perturbation would make the wing rock motion of a nondeterministic nature and an artificial mini-tip perturbation would make the wing rock motion deterministic. The artificial tip perturbation would, as its circumferential location is varied, generate three different types of motion patterns： （1） limit cycle oscillation, （2） irregular oscillation, （3） equilibrium state with tiny oscillation. The amplitude of rolling oscillation corresponding to the limit cycle oscillatory motion pattern is decreased with the wing location shifting downstream along the body axis.

Asymmetric flow effect in a horizontal natural ventilated tunnel with different aspect ratios under the influence of longitudinal fire locations

This paper has analyzed the asymmetric flow effect of fire-induced thermal flow in a horizontal tunnel under the natural ventilation condition by conducting large eddy simulations (LES). The key objective is to reveal and to have a better understanding of the asymmetric flow effect caused by the upstream and downstream tunnel length difference. The mechanism behind it can be explained based on the conservation of mass and dynamic force analysis on the smoke and fresh air. The strength of the asymmetric flow effect is characterized by the mass flow rate of the induced longitudinal flow (net mass flow rate of a cross-section). An empirical correlation to predict the induced longitudinal mass flow rate is proposed. Furthermore, the law of smoke and air flow distribution within a horizontal tunnel is established. The proportion of smoke (or air) flowing out (or coming in) through the opening increase (or decrease) linearly with the increasing distance between that opening to the fire location. The variation of the air flow with the longitudinal fire location in a tunnel is more sensitive than the smoke flow. Results have shown that as the fire approaches the tunnel exit from the middle of the tunnel, the smoke spilling out through this opening is reduced from 50% to 40%, while the fresh air incoming from this opening is increased from 50% to 100% and vice versa.

Unsteady cooperative flow in compression system

When there are several bodies with relative motion in a flow field,such as the flow in the compression system of modern aero-engine,the flow field will have certain special features,one of which is that the time-space structure of such multi-bodies unsteady vorticity flow field would be either of unsteady natural flow(UNF)pattern or of unsteady cooperative flow(UCF)pattern.If we further examine the aerodynamic design system of aero-engine,there is no mechanism for the unsteady cooperative flow to occur,in other words the flow field must be of the unsteady natural flow type.If certain technical measures can be adopted to transform UNF into UCF,the aerodynamic performances will surely be improved.This is the main task the author and their colleague have been devoted to and the results are reviewed in the present paper with emphases laid on basic ideas,technical approaches and experimental verifications.

Characteristics of the Asymmetric Flow of Tropical Cyclone Shanshan(2006) During Its Turning and Intensification Period

In this paper,characteristics of the asymmetric flow of Tropical Cyclone（TC） Shanshan（2006） during its turning and intensification period over the oceanic area east of Taiwan are investigated,based on the simulation results from the nonhydrostatic mesoscale model WRF（Weather Research and Forecasting）.It is found that the symmetric flow strengthened as the TC intensified,whereas the amplitude of the asymmetric flow of wavenumber 2 increased more significantly,which was strong enough to be comparable with or even exceed that of wavenumber 1,becoming the main part of the asymmetric flow sometimes.The asymmetric waves rotated around the TC center mainly counterclockwise.The closer to the center,the faster the asymmetric waves rotated.Moreover,the asymmetric flow rotated rapidly（slowly） during the slow（rapid） intensification of the TC,and the radial wavenumber showed an increase during the TC intensification.Furthermore,because of the superposition of intensified symmetric flow with the positive perturbation of the asymmetric flow,the maximum wind speed of TC Shanshan became larger.During the merger of the double eyewalls of Shanshan,the symmetric flow showed less increase in strength and the intensification of maximum wind speed was mainly related to the energy accumulation caused by the phase change of the asymmetric waves.The energy accumulation was realized when the asymmetric waves altered the strength and distribution of the inner and outer maximum wind cores,leading to the combination of the inner and outer eyewalls and eventually resulting in the intensification of the TC.

A novel architecture of electro-hydrostatic actuator with digital distribution

To address the flow imbalance rapidly of the asymmetric Electro-Hydrostatic Actuator(EHA),this paper presents a novel architecture of asymmetric EHA with the Digital Distribution(EHA-DD).It also improves the flow nonlinearity and control accuracy of the pump,especially in the low-speed condition.The digital distribution with two High-Speed on–off Valves(HSVs)not only balances flow instead of the check valves,but also replaces the pump at the low-speed to control output flow by adjusting the PWM signal.The pump and HSVs are the crucial components to control flow output.Firstly,the flow calculation models of the pump and the duty ratio inversed model of the HSV are obtained through experimental tests and the identification method.To get the control input signal for the required flow,the pump speed and PWM duty ratio for the HSVs are inversed to compensate for flow output.Further,a multimode digital flow distribution control method based on pump speed,mainly including the pump-controlled mode for large flow demand and valve-controlled mode for little flow demand,is proposed to control the accurate flow output actively.The step extension experiments based on the flow calculation models are conducted on the EHA-DD prototype under elastic,opposite varying load.The results demonstrate that the EHADD realizes little position error by accurate flow control,and it is also beneficial to improve the service life of the pump.

An experimental investigation on static directiona stability

A generic aircraft usually loses its static directional stability at moderate angle of attack(typically 20–30°). In this research, wind tunnel studies were performed using an aircraft model with moderate swept wing and a conventional vertical tail. The purpose of this study was to investigate flow mechanisms responsible for static directional stability. Measurements of force, surface pressure and spatial flow field were carried out for angles of attack from 0° to 46° and sideslip angles from-8° to 8°. Results of the wind tunnel experiments show that the vertical tail is the main contributor to static directional stability, while the fuselage is the main contributor to static directional instability of the model. In the sideslip attitude for moderate angles of attack, the fuselage vortex and the wing vortex merged together and changed asymmetrically as angle of attack increased on the windward side and leeward side of the vertical tail. The separated asymmetrical vortex flow around the vertical tail is the main reason for reduction in the static directional stability. Compared with the wing vortices, the fuselage vortices are more concentrated and closer to the vertical tail, so the yawing moment of vertical tail is more unstable than that when the wings are absent. On the other hand,the attached asymmetrical flow over the fuselage in sideslip leads to the static directional instability of the fuselage being exacerbated. It is mainly due to the predominant model contour blockage effect on the windward side flow over the model in sideslip, which is strongly affected by angle of attack.