The increased complexity and scales of international arbitration cases require a larger roster of righteous and experienced international arbitrators. The difficulty to define ethics or ethical obligation in the area ...The increased complexity and scales of international arbitration cases require a larger roster of righteous and experienced international arbitrators. The difficulty to define ethics or ethical obligation in the area of international arbitration has caused increased discussion and confusion. As a result,the tendency towards codification of ethical obligations would provide a more precise and practical mean to regulate arbitrators' behavior. The legal sources of ethical obligations of international arbitrators contain international conventions,national laws,arbitral institutions' rules as well as institutional codes of ethics. The main substantive ethical obligations include disclosing,investigating,conducting the arbitration in accordance with the arbitration agreement,maintaining confidentiality,being competent and diligent. Chinese authorities and foreign-related arbitral institutions should enhance the codification and uniformity of the international arbitrators' ethical obligations. The relevant strategies for revising the Arbitration Lawand arbitrators' ethical codes,unifying the foreign-related arbitration institution ethical rules and joining the harmonious progress of international arbitration should be considered correspondingly.展开更多
This paper considers the pricing problem of collateralized debt obligations tranches under a structural jump-diffusion model, where the asset value of each reference entity is generated by a geometric Brownian motion ...This paper considers the pricing problem of collateralized debt obligations tranches under a structural jump-diffusion model, where the asset value of each reference entity is generated by a geometric Brownian motion and jump with an asymmetric double exponential distribution. Conditioned on the common factor of individual entity, this paper gets the conditional distribution, and further obtains the loss distribution of the whole reference portfolio. Based on the semi-analytic approach, the fair spreads of collateralized debt obligations tranches, i.e., the prices of collateralized debt obligations tranches, are derived.展开更多
I Universal criminal jurisdiction,which is exercised by domestic courts of States,is considered today by some scholars as a panacea for fighting against impunity of perpetrators of the most heinous international crime...I Universal criminal jurisdiction,which is exercised by domestic courts of States,is considered today by some scholars as a panacea for fighting against impunity of perpetrators of the most heinous international crimes worldwide.展开更多
The extraterritorial application of international human rights conventions is the legal basis for states to regulate the extraterritorial human rights obligations of transnational corporations.In recent years,it has b...The extraterritorial application of international human rights conventions is the legal basis for states to regulate the extraterritorial human rights obligations of transnational corporations.In recent years,it has been increasingly challenged by the new trend of the United Nations governing the human rights responsibilities of transnational corporations.A study of existing international rules and practices shows that the extraterritorial application of international human rights conventions depends on the extraterritorial jurisdiction of the States Parties,which mainly includes the actual control over extraterritorial territories and individuals as well as the execution of authority outside the territories.In view of the above,if human rights violations by individuals or transnational corporations occur in areas outside their home territories or are authorized and orchestrated by the home state,the home state shall bear the corresponding human rights responsibilities.Given the restrictive scope of the abovementioned extraterritorial application of international human rights conventions,the international cooperation obligation and the"do no harm"principle may become a new legal basis for states to fulfill their extraterritorial human rights obligations in the future,thereby expanding the basis for the extraterritorial application of international human rights conventions to deal with more serious issues of human rights violations by transnational corporations.However,on the whole,the current identification and codification of states regulating the extraterritorial human rights obligations of transnational corporations should respect the existing rules of international human rights law and should not add any new human rights obligations on states.展开更多
The charterparty is a legal contract of employing a vessel. In shipping matters, it is a highly important document since it allocates obligations, rights, duties, liabilities, risks, earnings, costs and profits betwee...The charterparty is a legal contract of employing a vessel. In shipping matters, it is a highly important document since it allocates obligations, rights, duties, liabilities, risks, earnings, costs and profits between the contracted parties, namely, the shipowner and the charterer. The interpretation of the above mentioned matters, as well as the understanding of charterparty terms, is considered of critical importance in chartering practice. Therefore, this paper constitutes a review of the most important aspects arising from charterparties in the main types of charter. The present study is based on shipping practices followed in accordance with the English Common Law throughout the chartering process (pre-fixture, fixture, execution of the charter, post fixture). This is a synopsis about the distribution of the liabilities and expenses between the shipowner and the charterer in the most representative types of charter. The analysis is seen from a commercial stand point. Therefore, it is mostly addressed to the shipping practitioners, maritime economists, academics, students and researchers who seek to form a comprehensive view on the subject. It may also form a basis for further study on chartering aspects (legal, economic, managerial and practical).展开更多
The principal thesis of this paper is that one must grasp the importance, or even the centrality, of Hobbes's idea of commonwealth by institution in Leviathan in order to fully understand and appreciate his arguments...The principal thesis of this paper is that one must grasp the importance, or even the centrality, of Hobbes's idea of commonwealth by institution in Leviathan in order to fully understand and appreciate his arguments concerning the political obligation of obedience. Until recently, this idea has not been given its due significance in the literature. It has been commonly held that Hobbes's theorizing about the foundation of the state was based upon his concept of covenant. This paper argues that crucial aspects of political obligation generated by Hobbes's concept of covenant could not be understood without a clear understanding of the relationship among the three elements embodied in his idea of commonwealth by institution-covenant, authorization and right-transfer. While several attempts have been made on this topic, the pictures that have been drawn by Hobbesian scholars are more or less inadequate. In particular, the significant roles which authorization and right-transfer play in Hobbes's account of political obligation have yet been made clear. By shifting focus upon the idea of commonwealth by institution and taking it as Hobbes's mechanism of explaining the nature and origins of political obligation of obedience, this paper examines in detail the specific questions which Hobbes was addressing by coining this term and elicits what exactly he may have meant to convey to his audiences.展开更多
Are citizens morally obligated to obey and support their states? "Political Obligations and Authority" is perhaps A. John Simmons' most comprehensive summary of his mature views on this enduring philosophical prob...Are citizens morally obligated to obey and support their states? "Political Obligations and Authority" is perhaps A. John Simmons' most comprehensive summary of his mature views on this enduring philosophical problem. In this essay, Simmons critically engages Plato's dialogue Crito and culls from it three types of strategies for justifying political obligations: natural duty, associative, and transactional. Simmons argues that natural duty accounts are inherently incapable of providing moral grounding for political obligations, disqualifies both associative and transactional accounts on empirical grounds, and settles for a form of anarchism. I argue, assuming as Simmons does in this essay that natural duties imply obligations of support and obedience to political institutions, that the natural duty strategy promises to provide an escape route out of anarchism.展开更多
There is a transition from the objective laws or moral orders that precede human will in classical natural law to the subjective demands or rights emanating from human will in modern natural law,and it represents a hi...There is a transition from the objective laws or moral orders that precede human will in classical natural law to the subjective demands or rights emanating from human will in modern natural law,and it represents a historical debate on the shift from an obligation-based theory to a right-based theory.Strauss,within the context of this transition across time,assesses Thomas Hobbes's philosophy of law and recognizes him as the founder of modern natural rights theory.Using Wesley Newcomb Hohfeld's analysis of jurisprudence,Howard Warrender assesses the privilege nature of Hobbes's concept of natural rights and concludes that,since Hohfeld's privilege is the opposite of obligation and related to no-right,obligations cannot be derived from natural rights.Therefore,Warrender argues that Strauss's assertion requires correction.However,Warrender places excessive emphasis on Hohfeld's static separation of the concept of privilege within his theoretical system,overlooking the dynamic transformation from privilege to claim rights.In this regard,Hobbesian scholar Carlan's criticism of Warrender is valid.Meanwhile,Warrender's research holds theoretical significance in that he,under the premise of being a part of Hobbes'natural law tradition,transforms Hohfeld's flat,two-party legal rights relationships into a three-party legal rights structure,which could represent a potential innovation in the 20^(th)century legal philosophy.展开更多
This is a bioethical investigation into the nature of the endemic crisis,its survival,and the social construction of moral obligation during the Ebola crisis in Liberia,West Africa.The 2014 outbreak in West Africa was...This is a bioethical investigation into the nature of the endemic crisis,its survival,and the social construction of moral obligation during the Ebola crisis in Liberia,West Africa.The 2014 outbreak in West Africa was the most considerable,most severe,and most complex Ebola epidemic thus far.At the close of the crisis in Liberia,six thousand infected persons survived unexpectedly.The ethics of Ebola and survival is exceptionally complicated and requires a complex theoretical explanation.While a categorical analysis of ethical theory cannot cover the full scope of this moral dilemma,a single concept carried over a range of models does;and,that concept is known as moral obligation.Exploring the obligation of others toward Ebola survivors helps locate,justify,and analyze the fear-based system of morality that arose from the world’s most severe endemic crisis.Examining the obligations of government,community,individuals,and foreign research initiatives toward the survivors of the West African Ebola endemic in Monrovia,this article explores the varying moralities of endemic crisis culture as it examines the complexity of judgment related to social obligation.展开更多
“The Fundamental Rights and obligations of Citizens”, the title of Chapter II of the current Constitution of PRC, and the stipulation that citizens must fulfill certain obligations while enjoying rights have trigger...“The Fundamental Rights and obligations of Citizens”, the title of Chapter II of the current Constitution of PRC, and the stipulation that citizens must fulfill certain obligations while enjoying rights have triggered many debates. Considering the historical origin, constitutional philosophy, and the text and structure of the Constitution, the special provisions of the current Constitution are influenced by the principle of consistency of rights and obligations. The principle of consistency of rights and obligations in the Constitution is of complex connotation. Therefore, although the principle of consistency of rights and obligations effectively connects the public and private spheres, it ignores the diversity and differences of the interests and elements contained in the Constitution, the asymmetry of the normative status of fundamental rights and fundamental obligations,and the right of citizens to self-determination of personal interests.The principle of consistency of rights and obligations should be purposefully narrowed and concretized: In the context of public-private integration and risk society prevention, the principle of consistency of rights and obligations can be used as a supplement to the functional system of the Constitution;in the field of fundamental political obligations, the principle of consistency of rights and obligations should be in line with the requirements of the state to respect and protect human rights;in the field of fundamental social obligations, the exercise of fundamental rights by individuals is protected by the Constitution as long as they comply with the law and do not infringe upon the interests of the social community. The principle of the consistency of rights and obligations is only used as the negative constituents of the determination of rights and the basis for the effect against a third party of fundamental rights.展开更多
The case of Varif et al.v. Czech Republic concerned whether Czech legislation that imposed a moratorium on school attendance and a fine as a punitive mechanism for non-vaccination violates the European Convention on H...The case of Varif et al.v. Czech Republic concerned whether Czech legislation that imposed a moratorium on school attendance and a fine as a punitive mechanism for non-vaccination violates the European Convention on Human Rights by forcing children to receive routine vaccinations. In the ruling, the European Court of Human Rights reiterated that the choice of public health measures falls within the discretion of member states. Meanwhile, to limit the infringement of individual interests to what is necessary and reasonable, the European Court of Human Rights clarified the criteria for human rights protection for compulsory routine vaccination: It should have a legal basis, a lawful aim, and be in line with the “needs of a democratic society.” Accordingly, the European Court of Human Rights held that the legal provisions involved in the case did not violate the European Convention on Human Rights. As the first response of the European Court of Human Rights to compulsory vaccination, the case of Varif et al.v. Czech Republic strengthens the influence of the law of the European Union in the field of public health by upholding the discretion of member states. Besides, although the case involves only routine vaccination, the human rights protection criteria clarified in it can be taken as a reference for introducing vaccination measures against COVID-19 in the context of the global pandemic.展开更多
Mutualisms are cooperative interactions between members of different species. We focus on obligate mutualism, where each species cannot survive without the other. From a theoretical aspect, obligate mutualism is simil...Mutualisms are cooperative interactions between members of different species. We focus on obligate mutualism, where each species cannot survive without the other. From a theoretical aspect, obligate mutualism is similar to the relationship between male and female. Empirical data indicate a sex-ratio selection: male and female have a specific ratio in their population sizes. In the present paper, we apply lattice model to obligate mutualism between two species, and present a theory of “ratio selection” which is a generalization of sex-ratio selection. Computer simulations are carried out by two methods: local and global interactions. In the former, interactions occur between neighbouring cells, while in the latter they occur between any pair of cells. Simulations in both interactions show the so-called Allee effect: both species can survive, when both densities are large in some extent. However, we find a large difference between local and global simulations. In the case of local interaction, restriction for survival is found to be extremely severe compared to global interaction. Both species require a proper ratio for their sustainability. This result leads to the theory of ratio selection: when interaction occurs locally, the ratio of both species is uniquely determined. We discuss that the ratio selection explains not only the evolution of endosymbionts from free-living ancestors but also the evolution from endosymbionts to organelles.展开更多
With the knowledge that cuckoos and cowbirds lay their eggs parasitically,and that some hosts eject parasitic eggs,ornithologists began to ponder the question of how host females discriminate between a foreign egg and...With the knowledge that cuckoos and cowbirds lay their eggs parasitically,and that some hosts eject parasitic eggs,ornithologists began to ponder the question of how host females discriminate between a foreign egg and their own eggs,wondering how hosts 'know' which egg to remove.Results of one of the rst uncontrolled experiments were inappropriately interpreted to imply ejection was based on discordancy,with hosts simply ejecting the egg in the minority,or the 'odd-looking' egg.Controlled experiments eventually revealed that hosts rst learn the appearance of own their eggs and discriminate between them and any odd egg in their nest,regardless of which egg type is in the minority.Recent work has shown that discordancy may play a role in discrimination by males mated successively with females that lay polymorphic eggs.We examine the details of the early experiments,in light of recent advances in studies of egg recognition.An ability to recognize eggs also has been extended,implicitly,to include obligate brood parasites,as it underlies several hypotheses in explanation of the behavior of parasites toward their hosts.Egg recognition in parasites,however,has not been experimentally con rmed,nor has a mechanism been identi ed by which parasites could discriminate between their own eggs and the other eggs in a nest.We review hypotheses(parasite competition,egg removal and multiple parasitism,ma a,farming) that require the ability of obligate brood parasites to discriminate eggs at di erent levels and the potential mechanisms used by parasites to recognize their own eggs and suggest experiments to test for egg discrimination.An assessment of the egg recognition ability of parasites is germane to our understanding of how parasites counteract defenses of hosts.展开更多
基金a periodical achievement of China Scholarship Council National Construction of High-level University Postgraduate Program in2016(Liu Jin Fa No.3100).
文摘The increased complexity and scales of international arbitration cases require a larger roster of righteous and experienced international arbitrators. The difficulty to define ethics or ethical obligation in the area of international arbitration has caused increased discussion and confusion. As a result,the tendency towards codification of ethical obligations would provide a more precise and practical mean to regulate arbitrators' behavior. The legal sources of ethical obligations of international arbitrators contain international conventions,national laws,arbitral institutions' rules as well as institutional codes of ethics. The main substantive ethical obligations include disclosing,investigating,conducting the arbitration in accordance with the arbitration agreement,maintaining confidentiality,being competent and diligent. Chinese authorities and foreign-related arbitral institutions should enhance the codification and uniformity of the international arbitrators' ethical obligations. The relevant strategies for revising the Arbitration Lawand arbitrators' ethical codes,unifying the foreign-related arbitration institution ethical rules and joining the harmonious progress of international arbitration should be considered correspondingly.
基金Supported by the National Natural Science Foundation of China (70771018)the Natural Science Foundation of Shandong Province (2009ZRB019AV)Mathematical Subject Construction Funds and the Key Laboratory of Financial Information Engineering of Ludong University (2008)
文摘This paper considers the pricing problem of collateralized debt obligations tranches under a structural jump-diffusion model, where the asset value of each reference entity is generated by a geometric Brownian motion and jump with an asymmetric double exponential distribution. Conditioned on the common factor of individual entity, this paper gets the conditional distribution, and further obtains the loss distribution of the whole reference portfolio. Based on the semi-analytic approach, the fair spreads of collateralized debt obligations tranches, i.e., the prices of collateralized debt obligations tranches, are derived.
文摘I Universal criminal jurisdiction,which is exercised by domestic courts of States,is considered today by some scholars as a panacea for fighting against impunity of perpetrators of the most heinous international crimes worldwide.
基金the phased achievement of the CSHRS(China Society for Human Rights Studies)project“Human Rights Obligations of States Regulating the Extraterritorial Human Rights Obligations of Transnational Corporations”(CSHRS2020-25YB)
文摘The extraterritorial application of international human rights conventions is the legal basis for states to regulate the extraterritorial human rights obligations of transnational corporations.In recent years,it has been increasingly challenged by the new trend of the United Nations governing the human rights responsibilities of transnational corporations.A study of existing international rules and practices shows that the extraterritorial application of international human rights conventions depends on the extraterritorial jurisdiction of the States Parties,which mainly includes the actual control over extraterritorial territories and individuals as well as the execution of authority outside the territories.In view of the above,if human rights violations by individuals or transnational corporations occur in areas outside their home territories or are authorized and orchestrated by the home state,the home state shall bear the corresponding human rights responsibilities.Given the restrictive scope of the abovementioned extraterritorial application of international human rights conventions,the international cooperation obligation and the"do no harm"principle may become a new legal basis for states to fulfill their extraterritorial human rights obligations in the future,thereby expanding the basis for the extraterritorial application of international human rights conventions to deal with more serious issues of human rights violations by transnational corporations.However,on the whole,the current identification and codification of states regulating the extraterritorial human rights obligations of transnational corporations should respect the existing rules of international human rights law and should not add any new human rights obligations on states.
文摘The charterparty is a legal contract of employing a vessel. In shipping matters, it is a highly important document since it allocates obligations, rights, duties, liabilities, risks, earnings, costs and profits between the contracted parties, namely, the shipowner and the charterer. The interpretation of the above mentioned matters, as well as the understanding of charterparty terms, is considered of critical importance in chartering practice. Therefore, this paper constitutes a review of the most important aspects arising from charterparties in the main types of charter. The present study is based on shipping practices followed in accordance with the English Common Law throughout the chartering process (pre-fixture, fixture, execution of the charter, post fixture). This is a synopsis about the distribution of the liabilities and expenses between the shipowner and the charterer in the most representative types of charter. The analysis is seen from a commercial stand point. Therefore, it is mostly addressed to the shipping practitioners, maritime economists, academics, students and researchers who seek to form a comprehensive view on the subject. It may also form a basis for further study on chartering aspects (legal, economic, managerial and practical).
文摘The principal thesis of this paper is that one must grasp the importance, or even the centrality, of Hobbes's idea of commonwealth by institution in Leviathan in order to fully understand and appreciate his arguments concerning the political obligation of obedience. Until recently, this idea has not been given its due significance in the literature. It has been commonly held that Hobbes's theorizing about the foundation of the state was based upon his concept of covenant. This paper argues that crucial aspects of political obligation generated by Hobbes's concept of covenant could not be understood without a clear understanding of the relationship among the three elements embodied in his idea of commonwealth by institution-covenant, authorization and right-transfer. While several attempts have been made on this topic, the pictures that have been drawn by Hobbesian scholars are more or less inadequate. In particular, the significant roles which authorization and right-transfer play in Hobbes's account of political obligation have yet been made clear. By shifting focus upon the idea of commonwealth by institution and taking it as Hobbes's mechanism of explaining the nature and origins of political obligation of obedience, this paper examines in detail the specific questions which Hobbes was addressing by coining this term and elicits what exactly he may have meant to convey to his audiences.
文摘Are citizens morally obligated to obey and support their states? "Political Obligations and Authority" is perhaps A. John Simmons' most comprehensive summary of his mature views on this enduring philosophical problem. In this essay, Simmons critically engages Plato's dialogue Crito and culls from it three types of strategies for justifying political obligations: natural duty, associative, and transactional. Simmons argues that natural duty accounts are inherently incapable of providing moral grounding for political obligations, disqualifies both associative and transactional accounts on empirical grounds, and settles for a form of anarchism. I argue, assuming as Simmons does in this essay that natural duties imply obligations of support and obedience to political institutions, that the natural duty strategy promises to provide an escape route out of anarchism.
基金a phased project of two general projects:the Translation and Study of Hobbes’s Of Man(Project No.22YJA720009)a project of the Humanities and Social Sciences Planning Fund by the Ministry of Education+1 种基金the Research on Early Modern Western Sovereignty Theory(Project No.SK2022010)a project of the Basic Scientific Research of the Institutions of Higher-learning affiliated to Central Departments。
文摘There is a transition from the objective laws or moral orders that precede human will in classical natural law to the subjective demands or rights emanating from human will in modern natural law,and it represents a historical debate on the shift from an obligation-based theory to a right-based theory.Strauss,within the context of this transition across time,assesses Thomas Hobbes's philosophy of law and recognizes him as the founder of modern natural rights theory.Using Wesley Newcomb Hohfeld's analysis of jurisprudence,Howard Warrender assesses the privilege nature of Hobbes's concept of natural rights and concludes that,since Hohfeld's privilege is the opposite of obligation and related to no-right,obligations cannot be derived from natural rights.Therefore,Warrender argues that Strauss's assertion requires correction.However,Warrender places excessive emphasis on Hohfeld's static separation of the concept of privilege within his theoretical system,overlooking the dynamic transformation from privilege to claim rights.In this regard,Hobbesian scholar Carlan's criticism of Warrender is valid.Meanwhile,Warrender's research holds theoretical significance in that he,under the premise of being a part of Hobbes'natural law tradition,transforms Hohfeld's flat,two-party legal rights relationships into a three-party legal rights structure,which could represent a potential innovation in the 20^(th)century legal philosophy.
文摘This is a bioethical investigation into the nature of the endemic crisis,its survival,and the social construction of moral obligation during the Ebola crisis in Liberia,West Africa.The 2014 outbreak in West Africa was the most considerable,most severe,and most complex Ebola epidemic thus far.At the close of the crisis in Liberia,six thousand infected persons survived unexpectedly.The ethics of Ebola and survival is exceptionally complicated and requires a complex theoretical explanation.While a categorical analysis of ethical theory cannot cover the full scope of this moral dilemma,a single concept carried over a range of models does;and,that concept is known as moral obligation.Exploring the obligation of others toward Ebola survivors helps locate,justify,and analyze the fear-based system of morality that arose from the world’s most severe endemic crisis.Examining the obligations of government,community,individuals,and foreign research initiatives toward the survivors of the West African Ebola endemic in Monrovia,this article explores the varying moralities of endemic crisis culture as it examines the complexity of judgment related to social obligation.
文摘“The Fundamental Rights and obligations of Citizens”, the title of Chapter II of the current Constitution of PRC, and the stipulation that citizens must fulfill certain obligations while enjoying rights have triggered many debates. Considering the historical origin, constitutional philosophy, and the text and structure of the Constitution, the special provisions of the current Constitution are influenced by the principle of consistency of rights and obligations. The principle of consistency of rights and obligations in the Constitution is of complex connotation. Therefore, although the principle of consistency of rights and obligations effectively connects the public and private spheres, it ignores the diversity and differences of the interests and elements contained in the Constitution, the asymmetry of the normative status of fundamental rights and fundamental obligations,and the right of citizens to self-determination of personal interests.The principle of consistency of rights and obligations should be purposefully narrowed and concretized: In the context of public-private integration and risk society prevention, the principle of consistency of rights and obligations can be used as a supplement to the functional system of the Constitution;in the field of fundamental political obligations, the principle of consistency of rights and obligations should be in line with the requirements of the state to respect and protect human rights;in the field of fundamental social obligations, the exercise of fundamental rights by individuals is protected by the Constitution as long as they comply with the law and do not infringe upon the interests of the social community. The principle of the consistency of rights and obligations is only used as the negative constituents of the determination of rights and the basis for the effect against a third party of fundamental rights.
文摘The case of Varif et al.v. Czech Republic concerned whether Czech legislation that imposed a moratorium on school attendance and a fine as a punitive mechanism for non-vaccination violates the European Convention on Human Rights by forcing children to receive routine vaccinations. In the ruling, the European Court of Human Rights reiterated that the choice of public health measures falls within the discretion of member states. Meanwhile, to limit the infringement of individual interests to what is necessary and reasonable, the European Court of Human Rights clarified the criteria for human rights protection for compulsory routine vaccination: It should have a legal basis, a lawful aim, and be in line with the “needs of a democratic society.” Accordingly, the European Court of Human Rights held that the legal provisions involved in the case did not violate the European Convention on Human Rights. As the first response of the European Court of Human Rights to compulsory vaccination, the case of Varif et al.v. Czech Republic strengthens the influence of the law of the European Union in the field of public health by upholding the discretion of member states. Besides, although the case involves only routine vaccination, the human rights protection criteria clarified in it can be taken as a reference for introducing vaccination measures against COVID-19 in the context of the global pandemic.
文摘Mutualisms are cooperative interactions between members of different species. We focus on obligate mutualism, where each species cannot survive without the other. From a theoretical aspect, obligate mutualism is similar to the relationship between male and female. Empirical data indicate a sex-ratio selection: male and female have a specific ratio in their population sizes. In the present paper, we apply lattice model to obligate mutualism between two species, and present a theory of “ratio selection” which is a generalization of sex-ratio selection. Computer simulations are carried out by two methods: local and global interactions. In the former, interactions occur between neighbouring cells, while in the latter they occur between any pair of cells. Simulations in both interactions show the so-called Allee effect: both species can survive, when both densities are large in some extent. However, we find a large difference between local and global simulations. In the case of local interaction, restriction for survival is found to be extremely severe compared to global interaction. Both species require a proper ratio for their sustainability. This result leads to the theory of ratio selection: when interaction occurs locally, the ratio of both species is uniquely determined. We discuss that the ratio selection explains not only the evolution of endosymbionts from free-living ancestors but also the evolution from endosymbionts to organelles.
基金funded chiefly by grants from the Natural Sciences and Engineering Research Council of Canada
文摘With the knowledge that cuckoos and cowbirds lay their eggs parasitically,and that some hosts eject parasitic eggs,ornithologists began to ponder the question of how host females discriminate between a foreign egg and their own eggs,wondering how hosts 'know' which egg to remove.Results of one of the rst uncontrolled experiments were inappropriately interpreted to imply ejection was based on discordancy,with hosts simply ejecting the egg in the minority,or the 'odd-looking' egg.Controlled experiments eventually revealed that hosts rst learn the appearance of own their eggs and discriminate between them and any odd egg in their nest,regardless of which egg type is in the minority.Recent work has shown that discordancy may play a role in discrimination by males mated successively with females that lay polymorphic eggs.We examine the details of the early experiments,in light of recent advances in studies of egg recognition.An ability to recognize eggs also has been extended,implicitly,to include obligate brood parasites,as it underlies several hypotheses in explanation of the behavior of parasites toward their hosts.Egg recognition in parasites,however,has not been experimentally con rmed,nor has a mechanism been identi ed by which parasites could discriminate between their own eggs and the other eggs in a nest.We review hypotheses(parasite competition,egg removal and multiple parasitism,ma a,farming) that require the ability of obligate brood parasites to discriminate eggs at di erent levels and the potential mechanisms used by parasites to recognize their own eggs and suggest experiments to test for egg discrimination.An assessment of the egg recognition ability of parasites is germane to our understanding of how parasites counteract defenses of hosts.