The interaction between temperature and precipitation on the potential distribution range of Betula ermanii in the alpine treeline ecotone on the Changbai Mountain

Alpine treeline ecotones are highly sensitive to climate warming.The low temperature-determined alpine treeline is expected to shift upwards in response to global warming.However,little is known about how temperature interacts with other important factors to influence the distribution range of tree species within and beyond the alpine treeline ecotone.Hence,we used a GF-2 satellite image,along with bioclimatic and topographic variables,to develop an ensemble suitable habitat model based on the species distribution modeling algorithms in Biomod2.We investigated the distribution of suitable habitats for B.ermanii under three climate change scenarios(i.e.,low(SSP126),moderate(SSP370)and extreme(SSP585)future emission trajectories)between two consecutive time periods(i.e.,current-2055,and 2055-2085).By 2055,the potential distribution range of B.ermanii will expand under all three climate scenarios.The medium and high suitable areas will decline under SSP370 and SSP585scenarios from 2055 to 2085.Moreover,under the three climate scenarios,the uppermost altitudes of low suitable habitat will rise to 2,329 m a.s.l.,while the altitudes of medium and high suitable habitats will fall to 2,201 and2,051 m a.s.l.by 2085,respectively.Warming promotes the expansion of B.ermanii distribution range in Changbai Mountain,and this expansion will be modified by precipitation as climate warming continues.This interaction between temperature and precipitation plays a significant role in shaping the potential distribution range of B.ermanii in the alpine treeline ecotone.This study reveals the link between environmental factors,habitat distribution,and species distribution in the alpine treeline ecotone,providing valuable insights into the impacts of climate change on high-elevation vegetation,and contributing to mountain biodiversity conservation and sustainable development.

Seasonal snow cover patterns explain alpine treeline elevation better than temperature at regional scale

Unprecedented modern rates of warming are expected to advance alpine treelines to higher elevations,but global evidence suggests that current treeline dynamics are influenced by a variety of factors.Seasonal snow cover has an essential impact on tree recruitment and growth in alpine regions,which may in turn influence current treeline elevation;however,little research has been conducted on its role in regional treeline formation.Based on 11,804treeline locations in the eastern Himalayas,we extracted elevation,climate,and topographic data for treeline and snowline.Specifically,we used linear and structural equation modelling to assess the relationship between these environmental factors and treeline elevation,and the climate-snow-treeline interaction mechanism.The results showed that the treeline elevation increased with summer temperature and permanent or seasonal snowline elevation,but decreased with snow cover days and spring temperature at the treeline positions(P<0.001).Importantly,spring snowline elevation(33.4%)and seasonal snow cover days(21.1%)contributed the most to treeline elevation,outperforming the permanent snowline,temperature,precipitation,and light.Our results support the assertion that the temperature-moisture interaction affects treeline elevation in the eastern Himalayas,but we also found that the effects were strongly mediated by seasonal snow cover patterns.The increasing tendency of snow cover governed by climate humidification observed in the eastern Himalayas,is likely to limit future treeline advancement and may even cause treeline decline due to the mortality of the remaining old trees.Together,our findings highlight the role of seasonal snow cover patterns in determining treeline elevation in the eastern Himalayas,which should be considered when assessing the potential for treeline ascent in snow-mediated alpine systems elsewhere.

Indicator of climate variability:low treeline displacement in arid valleys of mountain areas,China

As climate change intensifies,finding an ecological indicator to quickly and accurately reflect the impact on mountain ecosystems is necessary.The low treeline/timberline,highly sensitive to climate variability and changes significantly within 5–10years,provides a new way to study the response to regional climate variability.This study explored the distribution and vertical displacement patterns of the low treeline in the Upper Minjiang River of China,using SPOT remote sensing images in 1999 and 2013and long-term positional observations.Using the Geodetector model,the study investigated the dominant climatic factors influencing the low treeline displacement.The results showed that the low treeline was located at 1700–3200 m elevation on sunny slopes(southeast,south,southwest,and west slopes)with slopes over 25°.From 1999 to 2013,the low treeline moved downward by 6 m from 2561±264m to 2555±265 m,along with a warm–humid climate tendency.The downward displacement was greater on slopes over 25°and shady slopes(-20 m and-10 m,respectively)than on slopes≤25°and sunny slopes.Additionally,the downward was greater in the warm and humid Zagunao River Basin(-15 m)compared to the arid valley center(-7 m)and the cold Heishui River Basin(-3 m).Meanwhile,the low treeline displacement correlated negatively with precipitation and relative humidity variations at the significance level of 0.05,with correlation coefficients of-0.572and-0.551,respectively.Variations in relative humidity and temperature significantly affected the spatial differentiation of low treeline displacement with influencing power of 0.246(p=0.036<0.05)and 0.183(p=0.032<0.05),respectively.Thus,the low treeline is a moisture-limited line,and its formation and variation are closely related to regional water–heat balance.The study clarifies the indicative value of the low treeline for climate variability in mountain areas and can provide references for ecological restoration in arid valleys.

Contemporary and Historic Population Structure of Abies spectabilis at Treeline in Barun Valley,Eastern Nepal Himalaya

Treeline ecotone dynamics of Abies spectabilis （D. Don） Mirb. in the Barun valley, Makalu Barun National Park, eastern Nepal Himalaya were studied by establishing seven plots （20 m x variable length） from the foresfline to the tree species limit： three plots on the south- and north-facing slopes each （S1-S3, N1-N3）, and one plot on the east- facing slope （E） in the relatively undisturbed forests. A dendroecological method was used to study treeline advance rate and recruitment pattern. In all the plots, most trees established in the early 20th century, and establishment in the second half of the 20th century was confined to the foresfline area. Treeline position has not advanced substantially in the Barun valley, with 0nly 22 m average elevational shift in the last 13o years, and with average current shifting rate of 14 cm/yr. Moreover, no significant relationship was found between tree age and elevation on the south-, north-, and east-facing slopes. The number of seedlings and saplings in near the treeline area was negligible compared to that near the foresfline area. Therefore, A. spectabilis treeline response to the temperature change was slow, despite the increasing temperature trend in the region. Beside the temperature change, factors such as high inter-annual variability in temperature, dense shrub cover, and local topography also play an important role in treeline advance and controlling recruitment pattern above the treeline.

A review of modern treeline migration, the factors controlling it and the implications for carbon storage

Numerous studies have reported that treelines are moving to higher elevations and higher latitudes.Most treelines are temperature limited and warmer climate expands the area in which trees are capable of growing.Hence,climate change has been assumed to be the main driver behind this treeline movement.The latest review of treeline studies was published in 2009 by Harsch et al.Since then,a plethora of papers have been published studying local treeline migration.Here we bring together this knowledge through a review of 142 treeline related publications,including 477 study locations.We summarize the information known about factors limiting tree-growth at and near treelines.Treeline migration is not only dependent on favorable growing conditions but also requires seedling establishment and survival above the current treeline.These conditions appear to have become favorable at many locations,particularly so in recent years.The review revealed that at 66%of these treeline sites forest cover had increased in elevational or latitudinal extent.The physical form of treelines influences how likely they are to migrate and can be used as an indicator when predicting future treeline movements.Our analysis also revealed that while a greater percentage of elevational treelines are moving,the latitudinal treelines are capable of moving at greater horizontal speed.This can potentially have substantial impacts on ecosystem carbon storage.To conclude the review,we present the three main hypotheses as to whether ecosystem carbon budgets will be reduced,increased or remain the same due to treeline migration.While the answer still remains under debate,we believe that all three hypotheses are likely to apply depending on the encroached ecosystem.Concerningly,evidence is emerging on how treeline migration may turn tundra landscapes from net sinks to net sources of carbon dioxide in the future.

Contribution of Mass Elevation Effect to the Altitudinal Distribution of Global Treelines

Alpine treeline, as a prominent ecological boundary between forested mountain slopes and alpine meadow/shrub, is highly complex in altitudinal distribution and sensitive to warming climate. Great efforts have been made to explore their distribution patterns and ecological mechanisms that determine these patterns for more than 100 years, and quite a number of geographical and ecophysiological models have been developed to correlate treeline altitude with latitude or a latitude related temperature. However,on a global scale, all of these models have great difficulties to accurately predict treeline elevation due to the extreme diversity of treeline site conditions.One of the major reasons is that "mass elevation effect"(MEE) has not been quantified globally and related with global treeline elevations although it has been observed and its effect on treeline elevations in the Eurasian continent and Northern Hemisphere recognized. In this study, we collected and compiled a total of 594 treeline sites all over the world from literatures, and explored how MEE affects globaltreeline elevation by developing a ternary linear regression model with intra-mountain base elevation(IMBE, as a proxy of MEE), latitude and continentality as independent variables. The results indicated that IMBE, latitude and continentality together could explain 92% of global treeline elevation variability, and that IMBE contributes the most(52.2%), latitude the second(40%) and continentality the least(7.8%) to the altitudinal distribution of global treelines. In the Northern Hemisphere, the three factors' contributions amount to 50.4%, 45.9% and 3.7% respectively; in the south hemisphere, their contributions are 38.3%, 53%, and 8.7%, respectively. This indicates that MEE, virtually the heating effect of macro-landforms, is actually the most significant factor for the altitudinal distribution of treelines across the globe, and that latitude is relatively more significant for treeline elevation in the Southern Hemisphere probably due to fewer macro-landforms there.

Positive adaptation of Salix eriostachya to warming in the treeline ecotone,East Tibetan Plateau

Understanding of treeline ecotone ecophysiological adaptation to climate warming is still very limited. Furthermore, it is difficult to predict which plant species could dominate in the future. For this reason, a study was conducted in the treeline ecotone, East Tibetan Plateau to detect the adaptation of the dwarf willow(Salix eriostachya) to experimental warming. Compared to ambient conditions, the experimental warming advanced the bud break by 12 days, delayed the leaf abscission by20 days, and prolonged the growing period by 28 days.It also increased photosynthesis(47%), number of leaves(333%), leaf area(310%), and carbon sequestration of the dwarf willow. Experimental warming did not affect carbon use efficiency, but decreased water use efficiency significantly.Experimental warming enhanced the clonal ramets of Salix eriostachya(+ 3.7 shrubs m-2). The frequent air temperature fluctuations had minor effect on Salix eriostachya. Based on these findings, we highlighted that Salix eriostachya could dominate in the community treeline ecotone of east Tibetan Plateau in the future climate warming scenario.

Decreasing nutrient concentrations in soils and trees with increasing elevation across a treelineecotone in Rolwaling Himal, Nepal

At a global scale, tree growth in alpine treeline ecotones is limited by low temperatures. At a local scale, however, tree growth at its upper limit depends on multiple interactions of influencing factors and mechanisms. The aim of our research was to understand local scale effects of soil properties and nutrient cycling on tree growth limitation, and their interactions with other abiotic and biotic factors in a near-natural Himalayan treeline ecotone. Soil samples of different soil horizons, litter, decomposition layers, and foliage samples of standing biomass were collected in four altitudinal zones along three slopes, and were analysed for exchangeable cations and nutrient concentrations, respectively. Additionally, soil and air temperature, soil moisture, precipitation, and tree physiognomy patterns were evaluated. Both soil nutrients and foliar macronutrient concentrations of nitrogen(N), magnesium(Mg), potassium(K), and foliar phosphorus(P) decrease significantly with elevation. Foliar manganese(Mn) concentrations, bycontrast, are extraordinarily high at high elevation sites. Potential constraining factors on tree growth were identified using multivariate statistical approaches. We propose that tree growth, treeline position and vegetation composition are affected by nutrient limitation, which in turn, is governed by low soil temperatures and influenced by soil moisture conditions.

Dendroclimatic response of Abies spectabilis at treeline ecotone of Barun Valley, eastern Nepal Himalaya

A dendroclimatic study was conducted in the treeline ecotone of Barun Valley, eastern Nepal, to determine the tree-ring climate response and ring width trend of Abies spectabilis. A 160-year-old chronology, from 1850 to 2010, was developed from 38 tree-ring samples. No higher growth in recent decades was observed in tree-ring width in this area. The mean temperature of the current year in February and in the combined winter months of December, January, and February showed significant positive correlation with tree-ring width, although no significant correlation was found between tree-ring width and the precipitation pattern of the region. This tree-ring climate response result is different from that in other studies in Nepal, which could be attributed to location and elevation.

Topographic Controls on Alpine Treeline Patterns on Changbai Mountain,China

【Title】【Author】【Addresses】1 The control mechanisms of topography on alpine treeline pattern are critical to understanding treeline dynamics and future changes. These mechanisms have not been understood quite well enough because of increasing human disturbance and low data resolution. In this study, the relationship between the treeline pattern and topography was analyzed based on high spatial resolution remote sensing data and a digital elevation model in an area in Changbai Mountain with little human disturbance. Future treeline patterns were also predicted. The results showed that （a） aspects with high solar radiation and low snow cover have a high coverage rate of trees, （b） the peak coverage rate of trees switches from low slopes （〈5°） to medium slopes （5°~25°） as the elevation rises because of the extreme environment, （c） the coverage rate of trees is a function that depends on environmental factors controlled by topography, （d） the future treeline pattern is controlled by new temperature mechanisms, new environmental factors and the reallocation effect of topography. Our research implies that topography controls the treeline pattern and changes in the treeline pattern associated with global warming, due to the effect of global warming on environmental factors. This study may well explain the causes of heterogeneous changes in the treeline pattern in the horizontal direction as well as differences in treeline response to climate warming.

Climate change evidence in tree growth and stand productivity at the upper treeline ecotone in the Polar Ural Mountains

Background:Recent warming is affecting species composition and species areal distribution of many regions.However,although most treeline studies have estimated the rates of forest expansion into tundra,still little is known about the long-term dynamic of stand productivity at the forest-tundra intersection.Here,we make use of tree-ring data from 350 larch(Larix sibirica Ledeb.)and spruce(Picea obovata Ledeb.)sampled along the singular altitudinal treeline ecotone at the Polar Urals to assess the dynamic of stand establishment and productivity,and link the results with meteorological observations to identify the main environmental drivers.Results:The analysis of stand instalment indicated that more than 90%of the living trees appeared after 1900.During this period,the stand became denser and moved 50m upward,while in recent decades the trees of both species grew faster.The maximum afforestation occurred in the last decades of the twentieth century,and the large number of encountered saplings indicates that the forest is still expanding.The upward shift coincided with a slight increase of May-August and nearly doubling of September-April precipitation while the increase in growth matched with an early growth season warming(June+0.27°C per decade since 1901).This increase in radial growth combined with the stand densification led to a 6-90 times increase of biomass since 1950.Conclusion:Tree-ring based twentieth century reconstruction at the treeline ecotone shows an ongoing forest densification and expansion accompanied by an increased growth.These changes are driven by climate change mechanism,whereby the leading factors are the significant increase in May-June temperatures and precipitation during the dormant period.Exploring of phytomass accumulation mechanisms within treeline ecotone is valuable for improving our understanding of carbon dynamics and the overall climate balance in current treeline ecosystems and for predicting how these will be altered by global change.

Contrasting changes in above- and below-ground biomass allocation across treeline ecotones in southeast Tibet

Under conditions of a warmer climate,the advance of the alpine treeline into alpine tundra has implications for carbon dynamics in mountain ecosystems.However,the above- and below-ground live biomass allocations among different vegetation types within the treeline ecotones are not well investigated.To determine the altitudinal patterns of above-/below-ground carbon allocation,we measured the root biomass and estimated the above-ground biomass(AGB) in a subalpine forest,treeline forest,alpine shrub,and alpine grassland along two elevational transects towards the alpine tundra in southeast Tibet.The AGB strongly declined with increasing elevation,which was associated with a decrease in the leaf area index and a consequent reduction in carbon gain.The fine root biomass(FRB) increased significantly more in the alpine shrub and grassland than in the treeline forest,whereas the coarse root biomass changed little with increasing altitudes,which led to a stable below-ground biomass(BGB) value across altitudes.Warm and infertile soil conditions might explain the large amount of FRB in alpine shrub and grassland.Consequently,the root toshoot biomass ratio increased sharply with altitude,which suggested a remarkable shift of biomass allocation to root systems near the alpine tundra.Our findings demonstrate contrasting changes in AGB and BGB allocations across treeline ecotones,which should be considered when estimating carbon dynamics with shifting treelines.

Remote sensing and geographic information systems techniques in studies on treeline ecotone dynamics

We performed a meta-analysis on over 100 studies applying remote sensing(RS)and geographic information systems(GIS)to understand treeline dynamics.A literature search was performed in multiple online databases,including Web of Knowledge(Thomson Reuters),Scopus(Elsevier),BASE(Bielefeld Academic Search Engine),CAB Direct,and Google Scholar using treeline-related queries.We found that RS and GIS use has steadily increased in treeline studies since 2000.Spatialresolution RS and satellite imaging techniques varied from low-resolution MODIS,moderate-resolution Landsat,to high-resolution WorldView and aerial orthophotos.Most papers published in the 1990s used low to moderate resolution sensors such as Landsat Multispectral Scanner and Thematic Mapper,or SPOT PAN(Panchromatic)and MX(Multispectral)RS images.Subsequently,we observed a rise in high-resolution satellite sensors such as ALOS,GeoEye,IKONOS,and WorldView for mapping current and potential treelines.Furthermore,we noticed a shift in emphasis of treeline studies over time:earlier reports focused on mapping treeline positions,whereas RS and GIS are now used to determine the factors that control treeline variation.

Response of tree-ring growth to climate at treeline ecotones in the Qilian Mountains,northwestern China

Climate constitutes the main limiting factor for tree-ring growth in high-elevation forests, and the relationship between tree-ring growth and climate is complex. Based on tree-ring chronology and meteorological data, the influence of precipitation, mean temperature and mean minimum temperature at yearly, seasonal and monthly scales on the tree-ring growth of Picea crossifolia was studied at treeline ecotones in the Qilian Mountains, northwestern China. The results show that growing season temperatures of previous and current years are important limiting factors on tree-ring growth, particularly June mean temperature and mean minimum temperature of current year. The precipitations in the previous winter and current spring have a positive correlation, and in the current fall has a negative correlation with tree-ring growth, but these correlations are not significant. Our results suggest that temperature controls tree-ring growth more strongly than precipitation at treeline ecotones in the Qilian Mountains.

Characteristics of timberline and treeline altitudinal distribution in Mt.Namjagbarwa and their geographical interpretation

Different types of vegetation patches are alternately and randomly distributed in a timberline ecotone where the upper limit is the treeline and the lower limit is the timberline.However,most studies on timberline/treeline altitudinal distributions have simplified timberline or treeline as continuous curves and disregarded the fuzziness of timberline/treeline and the randomness of different vegetation patch distributions in a timberline ecotone.To study the altitudinal distribution characteristics of timberline and treeline from the perspective of uncertainty theory,we constructed the timberline and treeline elevation cloud models in Mt.Namjagbarwa in east Himalayas.Subsequently,we established multiple linear regression models by using nine influencing factors,namely,aspect,slope,topographic relief,dryness index,average temperature in January and July,latitude,summit syndrome(represented by the vertical distance from the peak),and snow effect(represented by the nearest distance from the snow)as independent variables,and the elevations of timberline/treeline as dependent variables.Then we compared the contributions of the nine factors in timberline,treeline,and the core and peripheral areas of timberline and treeline.The results show that 1)the timberline/treeline elevation cloud model can represent the overall characteristics(especially the uncertainty)of the altitudinal distributions of the timberline/treeline well.The uncertainty of treeline’s altitudinal distribution is higher than that of timberline(entropy and hyper entropy:207.59 m and 70.36 m for treeline elevation cloud;entropy and hyper entropy:191.17 m and 50.13 m for timberline elevation cloud).2)The influence of climate and topography on timberline and treeline are similar.The average temperature in July has a significant negative correlation with the timberline/treeline elevation in Mt.Namjagbarwa,which is the most critical factor that affects timberline and treeline elevation,explaining the altitudinal distribution of 44.01%timberline and 46.74%treeline.However,the contributions of the nine factors in core and peripheral areas of timberline and treeline area are evidently different.

Topography and human disturbances are major controlling factors in treeline pattern at Barun and Manang area in the Nepal Himalaya

The alpine treeline ecotone is an important component of mountain ecosystems of the Nepal Himalaya; it plays a vital role in the livelihood of indigenous people,and provides ecosystem services. However,the region faces a problem of paucity of data on treeline characteristics at the regional and landscape scales. Therefore,we used Remote Sensing(RS),and Geographic Information Science(GIS) approaches to investigate cross-scale interactions in the treeline ecotone. Additionally,European Space Agency land cover map,International Center for Integrated Mountain Development(ICIMOD) land cover map,ecological map of Nepal,and United States Geological Survey Shuttle Radar Topography Mission-Digital Elevation Model were used to analyze treeline pattern at the regional scale. Digital Globe high-resolution satellite imagery of Barun(eastern Nepal) and Manang(central Nepal) were used to study treeline patterns at the landscape scale. Treeline elevation ranges from 3300-4300 m above sea level. Abies spectabilis,Betula utilis,and Pinus wallichiana are the main treeline-forming species in the Nepal Himalaya. There is an east to west treeline elevationgradient at the regional scale. No slope exposure is observed at the regional scale; however,at the landscape scale,slope exposure is present only in a disturbed area(Manang). Topography and human disturbance are the main treeline controlling factor in Barun and Manang respectively.

How does the stand structure of treeline-forming species shape the treeline ecotone in different regions of the Nepal Himalayas?

Stand structure dynamics are considered as major happenings in any forest as a response to environmental changes.However,this important topic is underrepresented in the treeline studies in the Nepal Himalayas.We aimed to investigate site-as well as species-specific changes in morphometric features(basal diameter,crown cover,density,and height)along the elevational gradient across treeline ecotones in response to recent environmental changes.The stand structure characteristics of Abies spectabilis,Pinus wallichiana,and Betula utilis across the treeline ecotone of three study sites in Eastern(Barun),Central(Manang),and Western(Dhorpatan)Nepal were analyzed to elucidate structural heterogeneities.Altogether,eight transects(20 m×(60–250 m))across the treeline ecotone were established.Trees of all life forms,trees(>2 m),saplings(0.5–2 m),and seedlings(<0.5 m),within each transect were enumerated and sampled for the morphometric features and age.Site-specific and species-specific stand structure dynamics were found.The rate of basal area increment was higher in Barun,but the Manang treeline,despite profound regeneration in recent years,had a low annual basal area increment.Moreover,the altitudinal distribution of age and morphometry were not consistent among those ecotones.Furthermore,intra-specific competition was not significant.The site-specific stand structure dynamics explain why treelines do not respond uniformly to increasing temperature.It invokes,in further studies,the incorporation of the tree’s morphometric adaptation traits,phenotypic plasticity,and interactions between species genotype and the environment.

Comparative analysis of the mass elevation effect and its implication for the treeline between the Tibetan and Bolivian plateaus based on solar radiation

As one of the main non-zonal factors,the mass elevation effect(MEE)has significant impacts on both regional climates and mountain ecological patterns.In recent years,with the development of quantitative techniques and methods,quantitative studies on the MEE and its implication on mountain altitudinal belts have developed rapidly.However,some issues have not been solved yet,such as high errors in spatial temperature estimations and difficulties in the definition and extraction of intramountain base elevation.Moreover,there is still a lack of comparative studies on the MEE and its influence on treelines and snowlines as most studies were conducted on specific mountains or plateaus.To compare the MEE magnitudes of the Tibetan Plateau(TP)and the Bolivian Plateau(BP),we estimated the correspondent air temperatures and simulated the solar radiations based on MODIS surface temperature,station observation,and treeline data.Then,we analyzed the elevation of the 10℃isotherms on the two plateaus,the temperatures at the same elevation,and the solar radiations.According to the mechanism of the MEE and the relationship of solar radiation and treeline,we constructed treeline models for the two plateaus through a stepwise regression analysis by considering several influencing factors of the MEE(e.g.,air temperature and precipitation)and using solar radiation as its proxy.The results showed that:(1)the MEE magnitude on the TP is equivalent to that on the BP although the former is slightly higher than the latter;(2)the MEE strongly influences the highest treelines in the northern and southern hemispheres,which both occur on the two plateaus.Notably,the treeline distribution models based on solar radiation had higher accuracies than those models with parameters of temperature and precipitation(the adjusted R^(2) values were 0.76 for the TP and 0.936 for the BP),indicating that solar radiation can be used to quantify the MEE and its implications on treelines.Overall,the results of this study can serve as a basis for subsequent analyses on the MEE’s impact factors.

Consequences of Future Expansion at the Arctic Treeline in Northernmost Norway

Seedlings from eight seed populations of mountain birch(Betula pubescens var.tortuosa),were transplanted to a site close to the town Vard?,in the Arctic part of the Varanger area in Northernmost Norway,in order to investigate the adaptation to climate change in different birch provenances and the implications for the treeline ecosystem and the local population.A comparable site was established at Kilpisj?rvi(500 masl)in Northern Finland close to treeline.Five replicates with 20 plants per replicate were established per site.The Vard?site was partly snow-free,sheltered by a willow thicket,and partly exposed on a ditched peat bog with thick snow cover from a nearby snowfence.Annual measurements were carried out on survival and growth parameters.The preliminary conclusion from this study is that local climate may be more important than the overall climatic variation in the adaptation and reforestation process in Northernmost Fennoscandia.In this process,the subarctic willow and shrub vegetation seems to be an important factor influencing the microclimate and seedling establishment.Species and provenances originating from areas with similar latitudes and climatic conditions as the reforestation area,were most successful.The Varanger area has always been a meeting place between different cultures,i.e.,the Sami,Norwegian,Finnish and Russian population.The study indicates that in a changed climate the birch forest area would expand because there will be more willow growth and consequently more safe sites for birch seedling establishment and growth,which would also create a better local climate for the human population.In this process local birch populations that are adapted to a more coastal climate,would have an advantage.Since birch has been shown to be an important resource for all these cultures,this would decrease the level of conflicts between the different groups of stakeholders about the resources in the area.

Dynamics of the Alpine Treeline Ecotone under Global Warming:A Review

The alpine treeline ecotone is defined as a forest-grassland or forest-tundra transition boundary either between subalpine forest and treeless grassland,or between subalpine forest and treeless tundra.The alpine treeline ecotone serves irreplaceable ecological functions and provides various ecosystem services.There are three lines associated with the alpine treeline ecotone,the tree species line(i.e.,the highest elevational limit of individual tree establishment and growth),the treeline(i.e.,the transition line between tree islands and isolated individual trees)and the timber line(i.e.,the upper boundary of the closed subalpine forest).The alpine treeline ecotone is the belt region between the tree species line and the timber line of the closed forest.The treeline is very sensitive to climate change and is often used as an indicator for the response of vegetation to global warming.However,there is currently no comprehensive review in the field of alpine treeline advance under global warming.Therefore,this review summarizes the literature and discusses the theoretical bases and challenges in the study of alpine treeline dynamics from the following four aspects:(1)Ecological functions and issues of treeline dynamics;(2)Methodology for monitoring treeline dynamics;(3)Treeline shifts in different climate zones;(4)Driving factors for treeline upward shifting.