基金financially supported by the‘Strategic Leading Science&Technology Program’of the Chinese Academy of Sciences(XDA23060604)the Chinese Academy of Sciences‘Light of West China’Program,the Second.Tibetan Plateau Scientific Expedition and Research(STEP)Program(2019QZKK0302)+1 种基金the National Natural Science Foundation of China(31770501)Key Laboratory of Ecology and Environment in Minority Areas(Minzu University of China),National Ethnic Affairs Commission(KLEEMA202103).