Polymorphism in egg coloration is prominent in the Common Cuckoo (Cuculus canorus) and a common host, the Ashy-throated Parrotbill (Paradoxornis alphonsianus). Egg polymorphism has probably evolved as a consequence of...Polymorphism in egg coloration is prominent in the Common Cuckoo (Cuculus canorus) and a common host, the Ashy-throated Parrotbill (Paradoxornis alphonsianus). Egg polymorphism has probably evolved as a consequence of frequency-dependent selection in both host and parasite, and has, according to human vision, resulted in discrete immaculate white, pale blue and blue egg phenotypes within a single population. However, egg mimicry assessment is not always straightforward, and previous studies have shown that human based comparisons applied to the coloration of bird eggs may be inadequate. Here, we objectively quantify egg color of both parasite and host by spectrophotometry and assess egg mimicry of the Common Cuckoo to the eggs of its Ashy-throated Parrotbill host. Our results revealed that egg reflectance spectra agree well with the assessment based on human vision that cuckoo eggs mimic those of the parrotbill host, in both visible (VIS) and ultraviolet (UV) ranges. However, the white cuckoo egg shows slightly poorer mimicry than the blue cuckoo egg in corresponding host clutches. We suggest that the white parrotbill egg morph (and subsequently the whitish cuckoo egg color) may have evolved after the evolution of the blue egg morph due to strong selection from parasites in the cuckoo-parrotbill system.展开更多
The principle of ocean wave spectrometers was first presented several decades ago to detect the directional wave spectrum with real-aperture radar(Jackson,1981). To invert wave spectra using an ocean wave spectrometer...The principle of ocean wave spectrometers was first presented several decades ago to detect the directional wave spectrum with real-aperture radar(Jackson,1981). To invert wave spectra using an ocean wave spectrometer,for simplicity,the hydrodynamic forcing and wave-wave interaction effect are neglected and a Gaussian slope probability density function(pdf) is used to calculate the normalized backscattering cross-section( σ 0) of the ocean surface. However,the real sea surface is non-Gaussian. It is not known whether the non-Gaussian property of the sea surface will affect the performance of the inversion of the wave spectrum if following existing inversion steps and methods. In this paper,the pdf of the sea surface slope is expressed as a Gram-Charlier fourth-order expansion,which is quasi-Gaussian. The modulation transfer function(MTF) is derived for a non-Gaussian slope pdf. The effects of non-Gaussian properties of the sea surface slope on the inversion process and result are then studied in a simulation of the SWIM(Surface Waves Investigation and Monitoring) instrument configuration to be used on the CFOSAT(China-France Oceanography Satellite) mission. The simulation results show that the mean trend of σ 0 depends on the sea slope pdf,and the peakedness and skewness coefficients of the slope pdf affect the shape of the mean trend of σ 0 versus incidence and azimuth; owing to high resolution of σ 0 in the range direction,MTF obtained using the mean trend of σ 0 is almost as accurate as that set in the direct simulation; in the inversion,if ignoring the non-Gaussian assumption,the inversion performances for the wave spectrum decrease,as seen for an increase in the energy error of the inverted wave slope spectrum. However,the peak wavelength and wave direction are the same for inversions that consider and ignore the non-Gaussian property.展开更多
The modal back-scattering matrix can be extracted from reverberation data. For high frequency cases the ’window smoothed’ processing has been proposed by E. C. Shang, T. F. Gao and D. J. Tang (2002) to extract the ...The modal back-scattering matrix can be extracted from reverberation data. For high frequency cases the ’window smoothed’ processing has been proposed by E. C. Shang, T. F. Gao and D. J. Tang (2002) to extract the ’window averaged’ back-scattering matrix. It is pointed out in this paper that in order to inverse the ’window averaged’ back-scattering matrix by changing the source depth data we have to assume that the matrix is not related to the source depth, and the numerical simulation on the question has been conducted.展开更多
When imaging ocean surface waves by X-band marine radar, the radar backscatter from the sea surface is modulated by the long surface gravity waves. The modulation transfer function (MTF) comprises tilt, hydrodynamic...When imaging ocean surface waves by X-band marine radar, the radar backscatter from the sea surface is modulated by the long surface gravity waves. The modulation transfer function (MTF) comprises tilt, hydrodynamic, and shadowing modulations. A conventional linear MTF was derived using HH-polarized radar observations under conditions of deep water. In this study, we propose a new quadratic polynomial MTF based on W-polarized radar measurements taken from heterogeneous nearshore wave fields. This new MTF is obtained using a radar-observed image spectrum and in situ buoy-measured wave frequency spectrum. We validate the MTF by comparing peak and mean wave periods retrieved from X-band marine radar image sequences with those measured by the buoy. It is shown that the retrieval accuracies of peak and mean wave periods of the new MTF are better than the conventional MTF. The results also show that the bias and root mean square errors of the peak and mean wave periods of the new MTF are 0.05 and 0.88 s, and 0.32 and 0.53 s, respectively, while those of the conventional MTF are 0.61 and 0.98 s, and 1.39 and 1.48 s, respectively. Moreover, it is also shown that the retrieval results are insensitive to the coefficients in the proposed MTF.展开更多
基金supported by the National Natural Science Foundation of China (Nos. 31071938 and 31272328 to WL, 31101646 and 31260514 to CY)Program for New Century Excellent Talents in University (NCET-10-0111 to WL)Key Project of Chinese Ministry of Education (No. 212136 to CY)
文摘Polymorphism in egg coloration is prominent in the Common Cuckoo (Cuculus canorus) and a common host, the Ashy-throated Parrotbill (Paradoxornis alphonsianus). Egg polymorphism has probably evolved as a consequence of frequency-dependent selection in both host and parasite, and has, according to human vision, resulted in discrete immaculate white, pale blue and blue egg phenotypes within a single population. However, egg mimicry assessment is not always straightforward, and previous studies have shown that human based comparisons applied to the coloration of bird eggs may be inadequate. Here, we objectively quantify egg color of both parasite and host by spectrophotometry and assess egg mimicry of the Common Cuckoo to the eggs of its Ashy-throated Parrotbill host. Our results revealed that egg reflectance spectra agree well with the assessment based on human vision that cuckoo eggs mimic those of the parrotbill host, in both visible (VIS) and ultraviolet (UV) ranges. However, the white cuckoo egg shows slightly poorer mimicry than the blue cuckoo egg in corresponding host clutches. We suggest that the white parrotbill egg morph (and subsequently the whitish cuckoo egg color) may have evolved after the evolution of the blue egg morph due to strong selection from parasites in the cuckoo-parrotbill system.
基金Supported by the National Science Foundation of China(No.40971185)the National High Technology Research and Development Program of China(863 Program)(No.2013AA09A505)
文摘The principle of ocean wave spectrometers was first presented several decades ago to detect the directional wave spectrum with real-aperture radar(Jackson,1981). To invert wave spectra using an ocean wave spectrometer,for simplicity,the hydrodynamic forcing and wave-wave interaction effect are neglected and a Gaussian slope probability density function(pdf) is used to calculate the normalized backscattering cross-section( σ 0) of the ocean surface. However,the real sea surface is non-Gaussian. It is not known whether the non-Gaussian property of the sea surface will affect the performance of the inversion of the wave spectrum if following existing inversion steps and methods. In this paper,the pdf of the sea surface slope is expressed as a Gram-Charlier fourth-order expansion,which is quasi-Gaussian. The modulation transfer function(MTF) is derived for a non-Gaussian slope pdf. The effects of non-Gaussian properties of the sea surface slope on the inversion process and result are then studied in a simulation of the SWIM(Surface Waves Investigation and Monitoring) instrument configuration to be used on the CFOSAT(China-France Oceanography Satellite) mission. The simulation results show that the mean trend of σ 0 depends on the sea slope pdf,and the peakedness and skewness coefficients of the slope pdf affect the shape of the mean trend of σ 0 versus incidence and azimuth; owing to high resolution of σ 0 in the range direction,MTF obtained using the mean trend of σ 0 is almost as accurate as that set in the direct simulation; in the inversion,if ignoring the non-Gaussian assumption,the inversion performances for the wave spectrum decrease,as seen for an increase in the energy error of the inverted wave slope spectrum. However,the peak wavelength and wave direction are the same for inversions that consider and ignore the non-Gaussian property.
文摘The modal back-scattering matrix can be extracted from reverberation data. For high frequency cases the ’window smoothed’ processing has been proposed by E. C. Shang, T. F. Gao and D. J. Tang (2002) to extract the ’window averaged’ back-scattering matrix. It is pointed out in this paper that in order to inverse the ’window averaged’ back-scattering matrix by changing the source depth data we have to assume that the matrix is not related to the source depth, and the numerical simulation on the question has been conducted.
基金Supported by the National High Technology Research and Development Program of China(863 Program)(No.2013AA09A505)the National Natural Science Foundation of China(Nos.41076119,41176160,41476158)+4 种基金the Priority Academic Program Development of Jiangsu Higher Education Institutions(PAPD)the Natural Science Youth Foundation of Jiangsu Province(No.BK2012467)the Natural Science State Key Foundation of Jiangsu Province(No.BK2011008)the National Natural Science Youth Foundation of China(No.41206171)the Startup Foundation for Introducing Talent of Nanjing University of Information Science and Technology(No.S8113078001)
文摘When imaging ocean surface waves by X-band marine radar, the radar backscatter from the sea surface is modulated by the long surface gravity waves. The modulation transfer function (MTF) comprises tilt, hydrodynamic, and shadowing modulations. A conventional linear MTF was derived using HH-polarized radar observations under conditions of deep water. In this study, we propose a new quadratic polynomial MTF based on W-polarized radar measurements taken from heterogeneous nearshore wave fields. This new MTF is obtained using a radar-observed image spectrum and in situ buoy-measured wave frequency spectrum. We validate the MTF by comparing peak and mean wave periods retrieved from X-band marine radar image sequences with those measured by the buoy. It is shown that the retrieval accuracies of peak and mean wave periods of the new MTF are better than the conventional MTF. The results also show that the bias and root mean square errors of the peak and mean wave periods of the new MTF are 0.05 and 0.88 s, and 0.32 and 0.53 s, respectively, while those of the conventional MTF are 0.61 and 0.98 s, and 1.39 and 1.48 s, respectively. Moreover, it is also shown that the retrieval results are insensitive to the coefficients in the proposed MTF.