基金supported by the National Key Research and Development Program of China[grant number 2016YFC1401803]the National Natural Science Foundation of China[grant numbers 41976019 and 42076020]+3 种基金the Strategic Priority Research Program of the Chinese Academy of Sciences[grant number XDA20060502]the open project of the State Key Laboratory of Tropical Oceanography,South China Sea Institute of Oceanology,Chinese Academy of Sciences[grant number LTO1910]the Research Program of the Southern Marine Science and Engineering Guangdong Laboratory(Guangzhou)[grant number GML2019ZD0306]the Key Research Program of the Chinese Academy of Sciences[grant number ZDRW-XH-2019-2]。