基金supported by the National Special Fund on Basic Research of Science and Technology of China(No.2014FY110100)supported by the National Special Fund on Basic Research of Science and Technology of China(No.2014FY110100)+5 种基金It is also partly sponsored by the Hunan Provincial Program for Development of Key Disciplines in Ecology(grant number No.0713)the National Natural Sciences Foundation of China(NSFC-30970327,31272271,31272272,31301861,31660609)Science and Technology Cooperation Project Foundation of Guizhou Province[grant number LH-2016/7303,LH-2017/7326],Natural Science Research Project Foundation of Guizhou Province of education[grant number KY[2018]345],Hunan Provincial Natural Science Foundation of China(No.11JJ1004/12JJ3028)the Program for New Century Excellent Talents in University(NCET-12-0717)China Postdoctoral Science Foundation(No.20100471221/201104506)the program of Hunan Provincial Science and Technology Plans(No.2010RS4006).