基金Supported by the National Natural Science Foundation of China(30370109,39899400,30499340)The Knowledge Innovation Program of Chinese Academy of Sciences(KSCX-SW-122,KSCX2-EW-J-28)+2 种基金Key Laboratory of Biodiversity and Biogeography,Kunming Institute of Botany,Chinese Academy of Sciences(KBB-200903)Doctoral Found of Bingzhou University(2010Y07)the Director Foundation of South China Botanical Garden,Chinese Academy of Sciences(2002-1112)