冬应肾而养藏

《黄帝内经》中明确提出＂五脏应四时,各有收受＂,具体说就是＂心者,生之本……为阳中之太阳,通于夏气;肺者,气之本……为阳中之太阴,通于秋气;肾者,主蛰,封藏之本……为阴中之少阴,通于冬气;肝者,罢极之本……为阳中之少阳,通于春气＂。这段话的意思是五脏各有对应的季节,肝对应春天、心对应夏天、肺对应秋天、肾对应冬天。所以,根据季节的不同,每一个脏器的养生调理侧重点也应该有所不同。

Biomass Allocation Patterns of Alpine Grassland Species and Functional Groups along a Precipitation Gradient on the Northern Tibetan Plateau

Variations in the fractions of biomass allocated to functional components are widely considered as plant responses to resource availability for grassland plants. Observations indicated shoots isometrically relates to roots at the community level but allometrically at the species level in Tibetan alpine grasslands. These differences may result from the specific complementarity of functional groups between functional components, such as leaf, root, stem and reproductive organ. To test the component complementary responses to regional moisture variation, we conducted a multi-site transect survey to measure plant individual size and component biomass fractions of common species belonging to the functional groups： forbs, grasses, legumes and sedges on the Northern Tibetan Plateau in peak growing season in 2010. Along the mean annual precipitation （MAP） gradient, we sampled 7o species, in which 2o are in alpine meadows, 20 in alpine steppes, 15 in alpine desert-steppes and 15 in alpine deserts, respectively. Our results showed that the size of alpine plants is small with individual biomass mostly lower than 1.0 g. Plants keep relative conservative component individual responses moisture functional fractions across alpine grasslands at the level. However, the complementary between functional components to variations specifically differ among groups. These results indicate that functional group diversity may be an effective tool for scaling biomass allocation patterns from individual up to community level. Therefore, it is necessary andvaluable to perform intensive and systematic studies on identification and differentiation the influences of compositional changes in functional groups on ecosystem primary services and processes.

Effects of Elevated Air Temperatures on Soil Thermal and Hydrologic Processes in the Active Layer in an Alpine Meadow Ecosystem of the Qinghai-Tibet Plateau

In this study,effects of elevated air temperatures on thermal and hydrologic process of the shallow soil in the active layer were investigated. Open-top chambers(OTCs)were utilized to increase air temperatures 1-2℃ in OTC-1 and 3-5℃ in OTC-2 in the alpine meadow ecosystem on the Qinghai- Tibetan Plateau.Results show that the annual air temperatures under OTC-1 and OTC-2 were 1.21℃ and 3.62℃ higher than the Control,respectively.The entirely-frozen period of shallow soil in the active layer was shortened and the fully thawed period was prolonged with temperature increase.The maximum penetration depth and duration of the negative isotherm during the entirely-frozen period decreased, and soil freezing was retarded in the local scope of the soil profile when temperature increased.Meanwhile, the positive isotherm during the fully-thawed period increased,and the soil thawing was accelerated.Soil moisture under different manipulations decreased with the temperature increase at the same depth. During the early freezing period and the early fully- thawed period,the maximum soil moisture under the Control manipulation was at 0.2 m deep,whereas under OTC-1 and OTC-2 manipulations,the maximum soil moisture were at 0.4-0.5 m deep. These results indicate that elevated temperatures led to a decrease of the moisture in the surface soil.The coupled relationship between soil temperature and moisture was significantly affected by the temperature increase.During the freezing and thawing processes, the soil temperature and moisture under different manipulations fit the regression model given by the equationθV=a/{1+exp[b(TS+c)]}+d.

Species Richness and Diversity of Alpine Grasslands on the Northern Tibetan Plateau:Effects of Grazing Exclusion and Growing Season Precipitation

Species richness and diversity indices （Shannon-Wiener index, Simpson dominance index and Pielou evenness index） in alpine grassland ecosystems （alpine meadow, alpine steppe and desert steppe） under grazing-excluded and freely grazed sites were investigated along the Northern Tibetan Plateau Alpine Grassland Transect during summer 2009 and 2010. We found that species richness and diversity have not been significantly altered by short-term grazing exclusion since 2006 at vegetation and regional scales. Species richness and diversity were mainly driven by growing season precipitation （GSP）, which accounted for over 87 % of the total variation observed, Species richness and diversity at grazing- excluded and freely grazed sites appear to respond to growing season precipitation in parallel. Species richness exponentially increased with GSP while diversity indices showed positively linear relationships with GSP. This indicates that GSP on the Northern Tibetan Plateau is crucial in regulating species richness and diversity and should be taken into account in future studies on alpine grassland conservation.