东北北部温带森林和干草地土壤养分分布及影响因素

土壤是陆地生态系统碳储存的重要场所,其养分变化与全球陆地碳循环密切相关。土壤养分是植物生长的重要保证,而土壤各养分之间是紧密联系的。理解土壤养分变化与环境因素的关系有助于更好地了解陆地生态系统碳、氮、磷循环。本研究以东北北部自东向西沿降水量梯度变化纬度带上的温带森林与干草地生态系统为研究对象,利用气象数据和野外土壤实测数据,分析了纬度带上不同植被类型土壤的有机碳、全氮、碳氮比、速效磷的空间分布格局及其与环境因子(年降水量、年均温、土壤pH值)的关系。研究纬度带上降水量自东向西逐渐减少,植被类型从温带森林过渡到干草原,与降水量和植被类型对应,植被生物量也自东向西呈现从高到低的分布梯度。研究结果表明:从整个研究带上来说,降水量与土壤pH值是土壤养分空间分布的决定因素,沿纬度带从东到西,随着降水量逐渐减少,土壤pH值逐渐增加,而土壤有机碳、全氮、碳氮比、速效磷含量逐渐减少。但如果将森林和草地分别讨论则发现,森林和草地生态系统的土壤养分环境控制因素有较大差别。对于草地生态系统而言,降水量和土壤pH值仍然是其土壤养分含量的控制因子,但森林生态系统由于所处区域降水量充足,降水量不再是其土壤养分的控制因子,降水量只与森林土壤碳氮比呈显著正相关。研究还发现森林土壤的速效磷含量与温度呈正相关,与土壤pH值呈负相关,说明温度对东北北部温带森林的土壤养分含量具有一定的控制作用。

4种多元聚合新策略在德国干草地类型划分中的应用

基于扩展了的Ｌａｎｃｅ和Ｗｉｌｉａｍｓ的多元聚合模型之上，现已提出了不少新的聚合策略。笔者用其中４种新策略，即新组内离差平方和法、新组内方差法、新组内平均距离法和加权平均距离法，对德国西北部Ｈｏｐｓｔｅｎ干草地进行了分类。结果表明，４种方法结果趋于一致，吻合度很高；所划分的类型与实际相符，生态意义明确，影响该草地分异的主要因素是土壤砂质含量和土壤保水能力，人类活动也是一重要影响因素。

攀世界荒漠戈壁科学之巅创中国极干草地治理之举——一个有成果有效益有活力的安西荒漠戈壁草地自然保护区通过阶段验收

1990年9月4～7日,国家农业部和环保局在安西县联合主持召开了安西荒漠戈壁草地自然保护区建设项目阶段验收会议。财政部、甘肃省及有关省区畜牧、环保、科研、大专院校的领导、专家、教授和代表共95人参加。会议由农业部畜牧兽医司副司长李仲昌和环保局同志主持。

Effects of Land Use Change on Ecosystem Services in Arid Area Ecological Migration

Ecological migration is the process of increasing the population density in the immigration area and transferring the ecological pressure from emigration area to immigration area. This process may result in significant changes in land use and land cover in the area of immigration and have an important effect on ecosystem services. Therefore, scientifically revealing the effects and differentiation mechanisms of ecological migration on ecosystem services is becoming an important issue related to the implementation of the national ecological migration strategy in China. This study employed the Hongsibu District as a typical example of ecological migration. Hongsibu District is located in the central Ningxia steppe and desert steppe areas. Remote sensing data covering five periods from the period before ecological migration in 1995 and after migration in 2000, 2005, 2010, and 2015 was used to measure the value of ecosystem services(ESV). A geographical detector model and the value of ecosystem services model were used to diagnose the dynamic mechanism of the effects of land use change on ecosystem services. The results showed that: 1) The development of large-scale ecological resettlement has caused the area of cultivated land and urbanized land area to increase significantly in the area of immigration, while the grass area decreased significantly. 2) The overall value of the Hongsibu ecosystem services increased in a form of a ‘V'. Among them, during the period of 1995–2005, the overall ESV decreased and had an annual rate of change of-0.67%. During the period of development 2005–2015, the ESV increased steadily, with an annual rate of change of 0.79%. 3) The proportion and total ESV in soil formation and protection, waste treatment, and biodiversity conservation of the Hongsibu District decreased from 57.61% in 1995 to 56.17% in 2015, indicating that the region's ecological regulation function slightly decreased. 4) The ESV in the Hongsibu District, showed a low distribution pattern of ecosystem services increasing from northeast to southwest, and the capacity of three townships, Hongsibu, Taiyangshan, and Liuquan, to provide ecosystem services gradually declined over time. The ecological service function of Xinzhuangji Township and Dahe Township gradually improved. 5) The sensitivity index of the ESV of each land use type was less than 1, indicating that the environment lacks flexibility in providing a strong ESV index in Hongsibu, which shows that the research results are reliable and believable. 6) During the study period, the decisive force of the change of land use on ecosystem services in Hongsibu District was: grassland(0.9934), climate regulation(0.9413), soil formation and protection(0.9321) and waste treatment(0.9241).

Drought shifts soil nematode trophic groups and mediates the heterotrophic respiration

As the most diverse metazoan taxa,soil nematodes serve a diversity of functions in soil food webs and thus can regulate microbial community composition and affect organic matter decomposition and nutrient turnover rates.Because nematodes depend on water flms to access food resources,drought can negatively affect nematode-microbial food webs,yet the impacts of drought on nematode diversity and abundance and how these changes may infuence food web members and their functions are hardly explored.Here,we coupled research along a drought gradient in arid and semiarid grasslands with a detailed intact plant-soil microcosm experiment to explore the patterns and mechanisms of how drought impacts nematode abundance and carbon footprint,microbial phospholipid fatty acid(PLFA)and heterotrophic soil respiration.Overall,in the feld and the microcosm experiments,we found that nematode abundance,carbon footprint and diversity,microbial PLFA and heterotrophic respiration were reduced under drier conditions.In addition,drought altered nematode and microbial community composition,through reducing the nematode channel ratio and increasing the relative fungivorous nematode abundance and the fungal to bacterial ratio.The soil decomposition channel shifted from a bacterial to a fungal pathway in response to drought,indicating decelerated heterotrophic respiration under drought.These results highlight the important contribution of soil nematodes and their associated microbial food web to soil carbon cycling.Our fndings underscore the need to incorporate key soil fauna into terrestrial ecosystem model evaluation.

Impact of Drought Stress on Net CO_2 Exchange above an Alpine Grassland Ecosystem in the Central Tibetan Plateau

Drought may impact the net ecosystem exchange of CO2 （NEE） between grassland ecosystems and the atmosphere during growth seasons. Here, carbon dioxide exchange and controlling factors in alpine grassland under drought stress in the hinterland of Tibetan Plateau （Damxung, Tibet, China） were investigated. Data were obtained using the covariance eddy technique in 2009. Severe drought stress appeared in the early growing season （May to early July） and September. Drought conditions during the early growing season limited grass production and the green leaf area index （GLAD increased slowly, with an obvious decline in June. When encountering severe water stress, diurnal patterns of NEE in the growth season altered with a peak carbon release around 16：00 h or a second carbon uptake period before sunset. NEE variations in daytime related most closely with O other than PAR when daily averaged @〈0.1 m3 m 3. Seasonal patterns of gross primary production （GPP） and NEE were also influenced by drought： the maximum and minimum of daily-integrated NEE were 0.9 g C m-2 d-1 on 3 July 2009, and -1.3 g C m-2 d-1 on 12 August 2009 with a GPP peak （-2.3 g C m-2 d-1） on the same day, respectively. Monthly NEE from May to July remained as carbon release and increased gradually; peak values of monthly NEE and GPP both appeared in August, but that of ecosystem respiration （R^co） was reached in July. Annual NEE, GPP and Reco of the alpine grassland ecosystem were 52.4, -158.1 and 210.5 g C m-2, respectively. Therefore, the grassland was a moderate source of COs to the atmosphere in this dry year. Interannual variation in NEE was likely related to different water conditions in the growing season. The three greatest contributors to seasonal variation in NEE, GPP and R^co respectively were GLAI〉Ta〉O, GLAI〉O〉PPT, and Ta〉GLAI〉PAR. Seasonality of GLAI explained 60.7% and 76.1% of seasonal variation in NEE and GPP, respectively. GPP or NEE was more sensitive than Reco to variation in GLAI, and ecosystem water conditions.

Influence of environmental factors on land-surface water and heat exchange during dry and wet periods in the growing season of semiarid grassland on the Loess Plateau

On the basis of information from the project ＂Land-surface Processes and their Experimental Study on the Chinese Loess Plateau＂, we analyzed differences in land-surface water and heat processes during the main dry and wet periods of the semiarid grassland growing season in Yuzhong County, as well as the influences of these environmental factors. Studies have shown that there are significant differences in changes of land-surface temperature and humidity during dry and wet periods. Daily average normalized temperature has an overall vertical distribution of ＂forward tilting＂ and ＂backward tilting＂ during dry and wet periods, respectively. During the dry period, shallow soil above 20-cm depth is the active temperature layer. The heat transfer rate in soil is obviously different during dry and wet periods. During the dry period, the ratio of sensible heat flux to net radiation （H/Rn） and the value of latent heat flux to net radiation （LE/Rn） have a linear relationship with 5-cm soil temperature; during the wet period, these have a nonlinear relationship with 5-cm soil temperature, and soil temperature of 16℃ is the critical temperature for changes in the land-surface water and heat exchange trend on a daily scale. During the dry period, H/Rn and LE/Rn have a linear relationship with soil water content. During the wet period, these have a nonlinear relationship with 5-cm soil water content, and 0.21 m^3 m^-3 is the critical point for changes in the land-surface water and heat exchange trend at daily scale. During the dry period, for vapor pressure deficit less than 0.7 kPa, H/Rn rises with increased vapor pressure deficit, whereas LEIRn decreases with that increase. When that deficit is greater than 0.7 kPa, both H/Rn and LE/Rn tend to be constant. During the wet period, H/Rn increases with the vapor pressure deficit, whereas LE/Rn decreases. The above characteristics directly reflect the effect of differences in land-surface environmental factors during land-surface water and heat exchange processes, and indirectly reflect the influences of cloud precipitation processes on those processes.

Effects of nitrogen addition and mowing on nitrogen-and water-use efficiency oi Artemisia frigida in a grassland restored from an abandoned cropland

Aims Competition among plants in a community usually depends on their nitrogen(N)-use efficiency(NUE)and water-use efficiency(WUE)in arid and semi-arid regions.Artemisia frigida is an indicator species in heavily degraded grassland,however,how its NUE and WUE respond to N addition in different successional stages is still unclear,especially with mowing,a common management practice in semi-arid grasslands.Methods Based on a long-term controlled experiment with N addition and mowing in an abandoned cropland from 2006 to 2013,we investigated the NUE and WUE oi A.frigida in two patches(i.e.grass and herb patches)in 2013 which represented two potential successional stages from herb to grass communities.The coverage of A.frigida was higher(about 50%)in the herb patch than in the grass patch(about 10%).Stable isotopic C(δ^(13)C)and N(δ^(15)N)as well as C and N pools were measured in plants and soils.NUE was calculated as leaf C/N,and leafδ^(13)C values were used as a proxy for WUE.Important Findings N addition did not affect WUE of A.frigida,but significantly decreased NUE by 42.9%and 26.6%in grass and herb patches,respectively.The response of NUE to N addition was related to altering utilization of different N sources(NH_(4)^(+)vs.NO_(3)^(-))by A.frigida according to the changed relationship between leafδ^(15)N/soil 615N and NUE.Mowing had no effect on NUE regardless of N addition,but significantly increased WUE by 2.3%for A.frigida without N addition in the grass patch.The addition of N reduced the positive effect of mowing on its WUE in grass patch.Our results suggested that decreased NUE and/or WUE of A.frigida under mowing and N addition could reduce its competition,and further accelerate restoration succession from the abandoned cropland to natural grassland in the semi-arid region.

Carbon dioxide exchange processes over the grassland ecosystems in semiarid areas of China

Based on the carbon fluxes measured over the grassland ecosystems in Inner Mongolia （UG79 site）, Loess Plateau （SACOL site）, and Tongyu, Jilin Province （TY site） in the semiarid areas from 2007 to 2008 with the eddy covariance method, we have investigated the carbon exchange processes over semiarid grassland ecosystem and its main affecting environmental variables. The precipitations at UG79 and TY sites in 2007 were below the historical average, especially for TY site, which was 50% be- low the historical average annual precipitation. The precipitation in SACOL site was close to average in 2007 but below average in 2008. The variation of monthly diurnal average NEE showed that the diurnal mean NEE decreased in the order of TY site, UG79 site, and SACOL site. However, a longer net carbon uptake period was observed at SACOL site. The diurnal course of NEE at UG79 site was similar between 2007 and 2008. The diurnal average NEE remained large during July and August in growing sea- son （May to September） at UG79 site, with maximum values approaching 0.08 mg C m^-2 s^-1 in August of 2008. The diurnal av- erage NEE of 2007 was larger than 2008 at SACOL site, with maximum values of 0.07 mg C m^-2 sq in September of 2007. A shorter carbon uptake period was recorded in 2007 at TY site, lasting from July to August. A larger diurnal average NEE oc- curred in 2008 at TY site, with maximum values of 0.12 mg C m^-2 s^-1. The ecosystem respirations of three sites were controlled by both soil temperature and soil volumetric water content （at a depth of 5 cm below the land surface）. Both UG79 site and SACOL site acted as a carbon sink during the growing periods of 2007 and 2008. Annual NEE in the growing seasons of 2007 and 2008 ranged from -68 to -50 g C m^-2 at UG79 site and from -109 to -55 g C m^-2at SACOL site. Alternation between car- bon source and carbon sink was found at TY site, with respective values of annual NEE in the growing seasons of 0.32 g C m^-2 and -73 g C m^-2 in 2007 and 2008. The magnitude and duration of carbon uptake depended mainly on the amount and timing of precipitation and the timing of the first effective rainfall during the growing season in semiarid grassland ecosystems.

Leaf and Ecosystem Gas Exchange Responses of Buffel Grass-Dominated Grassland to Summer Precipitation

Sporadic rain events that occur during summer play an important role in the initiation of biological activity of semi-arid grasslands.To understand how ecosystem processes of a buffel grass(Cenchrus ciliaris L.)-dominated grassland respond to summer rain events,an LI 6 400 gas exchange system was used to measure the leaf gas exchange and plant canopy chambers were used to measure net ecosystem CO2exchange(NEE) and ecosystem respiration(Reco), which were made sequentially during periods before rain(dry) and after rain(wet). Gross ecosystem photosynthesis(GEP) was estimated from NEE and Reco fluxes, and light use efficiency parameters were estimated using a rectangular hyperbola model. Prior to the monsoon rain, grassland biomass was non-green and dry exhibiting positive NEE(carbon source) and low GEP values during which the soil water became increasingly scarce. An initial rain pulse(60 mm) increased the NEE from pre-monsoon levels to negative NEE(carbon gain) with markedly higher GEP and increased green biomass. The leaf photosynthesis and leaf stomatal conductance were also improved substantially. The maximum net CO2uptake(i.e.,negative NEE) was sustained in the subsequent period due to multiple rain events. As a result, the grassland acted as a net carbon sink for 20 d after first rain. With cessation of rain(drying cycle), net CO2 uptake was reduced to lower values. High sensitivity of this grassland to rain suggests that any decrease in precipitation in summer may likely affect the carbon sequestration of the semiarid ecosystem.

Woody plant encroachment may decrease plant carbon storage in grasslands under future drier conditions

Aims Woody plants are widely distributed in various grassland types along the altitudinal/climatic gradients in Xinjiang,China.Considering previously reported change in carbon(C)storage following woody plant encroachment in grasslands and the mediating effect of climate on this change,we predicted that a positive effect of woody plants on plant C storage in semiarid grasslands may revert to a negative effect in arid grasslands.We first investigated the spatial variation of aboveground C(AGC)and belowground C(BGC)storage among grassland types and then tested our prediction.Methods We measured the living AGC storage,litter C(LC)and BGC storage of plants in two physiognomic types,wooded grasslands(aboveground biomass of woody plants at least 50%)and pure grasslands without woody plants in six grassland types representing a gradient form semiarid to arid conditions across Xinjiang.Important Findings Living AGC,LC,BGC and total plant C storage increased from desert to mountain meadows.These increases could also be explained by increasing mean annual precipitation(MAP)or decreasing mean annual temperature(MAT),suggesting that grassland types indeed represented an aridity gradient.Woody plants had an effect on the plant C storage both in size and in distribution relative to pure grasslands.The direction and strength of the effect of woody plants varied with grassland types due to the mediating effect of the climate,with wetter conditions promoting a positive effect of woody plants.Woody plants increased vegetation-level AGC through their high AGC relative to herbaceous plants.However,more negative effects of woody plants on herbaceous plants with increasing aridity led to a weaker increase in the living AGC in arid desert,steppe desert and desert steppe than in the less arid other grassland types.Under greater aridity(lower MAP and higher MAT),woody plants allocated less biomass to roots and had lower BGC and had a more negative impact on herbaceous plant production,thereby reducing vegetation-level BGC in the desert,steppe desert and desert steppe.In sum,this resulted in a negative effect of woody plants on total plant C storage in the most arid grasslands in Xinjiang.As a consequence,we predict that woody plant encroachment may decrease rather than increase C storage in grasslands under future drier conditions.