基金supported by grants from the National Natural Science Foundation of China(41731175 and 41988101)the Strategic Priority Research Program A of the Chinese Academy of Sciences(XDA20050101)+1 种基金the Joint Key Research Fund under a cooperative agreement between the National Natural Science Foundation of China(NSFC)and Tibet Autonomous Region(TAR)(U20A2005)the Second Tibetan Plateau Scientific Expedition and Research(STEP)program(2019QZKK0608 and 2019QZKK0302).